Citation:

PANGAEA.995107 (DOI registration in progress)

Name: Seawater carbonate chemistry and ocean alkalinity enhancement on Oikopleura dioica abundances and their house production capacity
Published: 2026-05-19
License: https://creativecommons.org/licenses/by/4.0/
Keywords: Animalia; Behaviour; Biomass/Abundance/Elemental composition; Chordata; Coast and continental shelf; Growth/Morphology; Laboratory experiment; Mesocosm or benthocosm; North Atlantic; Ocean alkalinity enhancement; Oikopleura dioica; Other studied parameter or process; Pelagos; Single species; Temperate; Zooplankton

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Published: 2026-05-19

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Abstract:

Here, we assessed the response of a key gelatinous zooplankton species to ocean alkalinity enhancement in a 53-day mesocosm experiment in a temperate Norwegian fjord. Oikopleura dioica is a globally distributed zooplankton member, known for its high secondary production capacity and key role in vertical carbon flux. O. dioica continuously produces mucous feeding structures ('houses'), which efficiently retain submicron particles. Once discarded, these houses can sink rapidly and contribute to vertical carbon exports. To test the impacts of OAE on O. dioica abundances and their house production capacity, we exposed natural plankton communities to non-CO2-equilibrated OAE scenarios spanning a ΔTA range from 0–600 μmol/kg, using silicate-based (olivine) and calcium-based (slaked lime) minerals. Population dynamics of O. dioica were monitored alongside the plankton community, and targeted bottle incubations were used to quantify house production and feeding rates.

This dataset is included in the OA-ICC data compilation maintained in the framework of the IAEA Ocean Acidification International Coordination Centre (see https://oa-icc.ipsl.fr). Original data were downloaded from zenodo (see Source) by the OA-ICC data curator. In order to allow full comparability with other ocean acidification data sets, the R package seacarb (Gattuso et al, 2024) was used to compute a complete and consistent set of carbonate system variables, as described by Nisumaa et al. (2010). In this dataset the original values were archived in addition with the recalculated parameters (see related PI). The date of carbonate chemistry calculation by seacarb is 2026-05-18.

Keyword(s):

Animalia; Behaviour; Biomass/Abundance/Elemental composition; Chordata; Coast and continental shelf; Growth/Morphology; Laboratory experiment; Mesocosm or benthocosm; North Atlantic; Ocean alkalinity enhancement; Oikopleura dioica; Other studied parameter or process; Pelagos; Single species; Temperate; Zooplankton

Related to:

Bhaumik, Amrita; Sánchez, Nicolás; Goldenberg, Silvan Urs; Spjelkavik, Synne; Couret, María; Riebesell, Ulf; Boersma, Maarten; Jaspers, Cornelia (2026): Resilience of the gelatinous zooplankton species Oikopleura dioica to ocean alkalinity enhancement. PLoS ONE, 21(3), e0344503, https://doi.org/10.1371/journal.pone.0344503

Schneider, Julieta; Riebesell, Ulf; Moras, Charly; Marín-Samper, Laura; Kittu, Leila; Ortiz Cortes, Joaquin; Schulz, Kai Georg (2025): KOSMOS 2022 Bergen mesocosm study on ocean alkalinity enhancement: carbonate chemistry , CO2 flux data and calcification estimates [dataset]. PANGAEA, https://doi.org/10.1594/PANGAEA.983061 (The original carbonate chemistry data as well as temperature and salinity data were downloaded from this dataset.)

Source:

Bhaumik, Amrita; Sánchez, Nicolás; Goldenberg, Silvan Urs; Spjelkavik, Synne; Couret, Maria; Riebesell, Ulf; Boersma, Maarten; Jaspers, Cornelia (2025): Data underlying figures for: Resilience of the gelatinous zooplankton species Oikopleura dioica to ocean alkalinity enhancement (PONE-D-25-40260) [dataset]. Zenodo, https://doi.org/10.5281/ZENODO.18011790

References:

Nisumaa, Anne-Marin; Pesant, Stephane; Bellerby, Richard G J; Delille, Bruno; Middelburg, Jack J; Orr, James C; Riebesell, Ulf; Tyrrell, Toby; Wolf-Gladrow, Dieter A; Gattuso, Jean-Pierre (2010): EPOCA/EUR-OCEANS data compilation on the biological and biogeochemical responses to ocean acidification. Earth System Science Data, 2(2), 167-175, https://doi.org/10.5194/essd-2-167-2010

Yang, Yan; Brockmann, Patrick; Galdino, Carolina; Schindler, Uwe; Gazeau, Frédéric (2024): An update of data compilation on the biological response to ocean acidification and overview of the OA-ICC data portal. Earth System Science Data, 16(8), 3771-3780, https://doi.org/10.5194/essd-16-3771-2024

Yang, Yan; Hansson, L; Gattuso, Jean-Pierre (2016): Data compilation on the biological response to ocean acidification: an update. Earth System Science Data, 8(1), 79-87, https://doi.org/10.5194/essd-8-79-2016

Related code / software:

Gattuso, Jean-Pierre; Epitalon, Jean-Marie; Lavigne, Héloïse; Orr, James; Gentili, Bernard; Hagens, Mathilde; Hofmann, Andreas; Mueller, Jens-Daniel; Proye, Aurélien; Rae, James; Soetaert, Karline (2024): seacarb: seawater carbonate chemistry with R. R package version 3.3.3. https://cran.r-project.org/web/packages/seacarb/index.html

Project(s):

Ocean Acidification International Coordination Centre (OA-ICC)

Parameter(s):

#	Name	Short Name	Unit	Principal Investigator	Method/Device	Comment
1	Type of study	Study type		Bhaumik, Amrita
2	Species, unique identification	Species UID		Bhaumik, Amrita
3	Species, unique identification	Species UID		Bhaumik, Amrita
4	Species, unique identification	Species UID		Bhaumik, Amrita
5	Day of experiment	DOE	day	Bhaumik, Amrita
6	Phase	Phase		Bhaumik, Amrita
7	Treatment	Treat		Bhaumik, Amrita		Mineral_type
8	Change in alkalinity, total	AT chn	µmol/kg	Bhaumik, Amrita
9	Abundance per volume	Abund v	#/m³	Bhaumik, Amrita
10	Abundance	Abund	%	Bhaumik, Amrita
11	Abundance per volume	Abund v	#/m³	Bhaumik, Amrita		small size
12	Abundance per volume	Abund v	#/m³	Bhaumik, Amrita		intermediate size
13	Abundance per volume	Abund v	#/m³	Bhaumik, Amrita		large size
14	House production rate, per individual	House prod/ind	#/#/day	Bhaumik, Amrita
15	Clearance rate per individual	CR	ml/#/h	Bhaumik, Amrita
16	Salinity	Sal		Bhaumik, Amrita
17	Temperature, water	Temp	°C	Bhaumik, Amrita
18	Alkalinity, total	AT	µmol/kg	Bhaumik, Amrita
19	pH, total scale	pHT		Bhaumik, Amrita	Potentiometric	T_25
20	pH, total scale	pHT		Bhaumik, Amrita	Potentiometric	T_25 correct
21	pH, total scale	pHT		Bhaumik, Amrita	Calculated using CO2SYS	T_insitu
22	Carbon, inorganic, dissolved	DIC	µmol/kg	Bhaumik, Amrita	Calculated using CO2SYS
23	Partial pressure of carbon dioxide (water) at sea surface temperature (wet air)	pCO2water_SST_wet	µatm	Bhaumik, Amrita	Calculated using CO2SYS
24	Calcite saturation state	Omega Cal		Bhaumik, Amrita	Calculated using CO2SYS
25	Aragonite saturation state	Omega Arg		Bhaumik, Amrita	Calculated using CO2SYS
26	Bicarbonate ion	[HCO3]-	µmol/kg	Bhaumik, Amrita	Calculated using CO2SYS
27	Carbon dioxide	CO2	µmol/kg	Bhaumik, Amrita	Calculated using CO2SYS
28	Carbonate system computation flag	CSC flag		Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
29	Carbon dioxide	CO2	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
30	Fugacity of carbon dioxide (water) at sea surface temperature (wet air)	fCO2water_SST_wet	µatm	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
31	Partial pressure of carbon dioxide (water) at sea surface temperature (wet air)	pCO2water_SST_wet	µatm	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
32	Bicarbonate ion	[HCO3]-	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
33	Carbonate ion	[CO3]2-	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
34	Carbon, inorganic, dissolved	DIC	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
35	Aragonite saturation state	Omega Arg		Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
36	Calcite saturation state	Omega Cal		Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)

License:

Creative Commons Attribution 4.0 International (CC-BY-4.0)

Status:

Curation Level: Enhanced curation (CurationLevelC)

Size:

2860 data points

Data

Download dataset as tab-delimited text — use the following character encoding:

1 Study type	2 Species UID	3 Species UID	4 Species UID	5 DOE [day]	6 Phase	7 Treat (Mineral_type)	8 AT chn [µmol/kg]	9 Abund v [#/m³]	10 Abund [%]	11 Abund v [#/m³] (small size)	12 Abund v [#/m³] (intermediate size)	13 Abund v [#/m³] (large size)	14 House prod/ind [#/#/day]	15 CR [ml/#/h]	16 Sal	17 Temp [°C]	18 AT [µmol/kg]	19 pHT (T_25, Potentiometric)	20 pHT (T_25 correct, Potentiometric)	21 pHT (T_insitu, Calculated using CO...)	22 DIC [µmol/kg] (Calculated using CO2SYS)	23 pCO2water_SST_wet [µatm] (Calculated using CO2SYS)	24 Omega Cal (Calculated using CO2SYS)	25 Omega Arg (Calculated using CO2SYS)	26 [HCO3]- [µmol/kg] (Calculated using CO2SYS)	27 CO2 [µmol/kg] (Calculated using CO2SYS)	28 CSC flag (Calculated using seacarb afte...)	29 CO2 [µmol/kg] (Calculated using seacarb afte...)	30 fCO2water_SST_wet [µatm] (Calculated using seacarb afte...)	31 pCO2water_SST_wet [µatm] (Calculated using seacarb afte...)	32 [HCO3]- [µmol/kg] (Calculated using seacarb afte...)	33 [CO3]2- [µmol/kg] (Calculated using seacarb afte...)	34 DIC [µmol/kg] (Calculated using seacarb afte...)	35 Omega Arg (Calculated using seacarb afte...)	36 Omega Cal (Calculated using seacarb afte...)
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Ca	0	1521.96	4.47	244.53	876.92	400.52			32.810	9.60	2210.0	7.861	7.860	8.098	2027.29	347.0	3.17	2.01	1880.3	15.57	8	15.16	336.76	338.07	1874.53	134.37	2024.07	2.05	3.24
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Ca	150	1391.27	3.90	193.93	784.17	413.16			32.740	9.62	2346.3	8.089	8.089	8.334	2034.81	194.0	5.25	3.32	1808.6	8.70	8	8.46	187.85	188.58	1799.79	221.95	2030.20	3.39	5.36
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Ca	300	1762.27	5.57	244.53	952.81	564.94			32.770	9.62	2497.6	8.276	8.290	8.540	2030.65	113.2	7.93	5.02	1697.2	5.08	8	4.93	109.47	109.90	1685.59	334.27	2024.79	5.11	8.07
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Ca	450	1933.02	5.34	413.16	1037.12	482.73			32.770	9.65	2608.3	8.380	8.401	8.654	2034.66	83.4	9.85	6.24	1622.8	3.73	8	3.61	80.37	80.68	1608.61	415.25	2027.48	6.35	10.03
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Ca	600	1266.89	4.59	109.61	834.76	322.52			32.820	9.64	2734.8	8.483	8.511	8.767	2039.63	60.8	12.10	7.66	1535.4	2.72	8	2.63	58.52	58.75	1519.80	509.33	2031.76	7.78	12.29
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Ca	0	1439.75	3.67	126.48	860.05	453.22			32.806	10.28	2199.6	7.856	7.856	8.083	2019.27	359.1	3.14	1.99	1873.4	15.75	8	15.37	349.13	350.48	1867.92	132.85	2016.15	2.03	3.21
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Ca	150	1041.34	2.71	31.62	581.80	427.92			32.755	10.30	2340.8	8.086	8.086	8.319	2031.57	200.8	5.22	3.31	1806.5	8.80	8	8.60	195.32	196.07	1798.75	220.18	2027.53	3.37	5.32
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Ca	300	1075.07	2.85	52.70	581.80	440.57			32.794	10.29	2491.2	8.270	8.284	8.522	2029.77	118.5	7.84	4.97	1699.8	5.19	8	5.06	114.91	115.35	1689.12	330.15	2024.33	5.05	7.97
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Ca	450	1406.02	4.21	94.86	767.30	543.86			32.787	10.30	2605.1	8.375	8.396	8.637	2036.34	87.1	9.78	6.20	1627.5	3.82	8	3.71	84.30	84.63	1614.77	411.39	2029.87	6.29	9.93
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Ca	600	883.25	2.85	63.24	463.76	356.25			32.834	10.31	2732.9	8.482	8.510	8.754	2038.92	62.8	12.09	7.66	1535.2	2.75	8	2.67	60.65	60.88	1521.00	508.08	2031.75	7.77	12.26
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Ca	0	820.56	2.13	126.48	487.50	206.58			32.827	10.11	2214.3	7.853	7.853	8.082	2034.39	362.0	3.14	1.99	1888.2	15.96	8	15.60	352.31	353.67	1883.09	132.80	2031.49	2.03	3.21
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Ca	150	956.49	2.68	137.02	619.22	200.26			32.778	10.12	2341.1	8.077	8.077	8.313	2036.99	204.6	5.14	3.25	1815.2	9.02	8	8.80	198.69	199.46	1807.27	216.77	2032.84	3.32	5.23
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Ca	300	895.89	2.39	305.66	421.60	168.64			32.804	10.12	2494.6	8.263	8.276	8.517	2038.27	120.5	7.75	4.91	1711.7	5.31	8	5.17	116.91	117.37	1701.18	326.59	2032.95	4.99	7.88
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Ca	450	558.61	1.51	126.48	263.50	168.64			32.790	10.14	2607.5	8.365	8.385	8.629	2047.36	89.6	9.63	6.10	1644.4	3.95	8	3.83	86.65	86.98	1631.33	405.50	2040.66	6.20	9.79
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Ca	600	463.76	1.18	52.70	263.50	147.56			32.828	10.14	2734.2	8.476	8.504	8.750	2045.99	63.7	11.99	7.60	1546.4	2.81	8	2.72	61.60	61.83	1532.52	503.79	2039.03	7.70	12.16
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Ca	0	1193.12	7.87	168.64	600.77	423.70			32.846	10.04	2204.5	7.847	7.847	8.077	2027.64	365.2	3.09	1.96	1883.5	16.14	8	15.76	355.27	356.65	1878.31	130.64	2024.71	2.00	3.15
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Ca	150	1618.93	5.51	158.10	853.73	607.10			32.781	10.06	2341.1	8.068	8.068	8.305	2042.29	209.3	5.05	3.20	1823.8	9.25	8	9.01	203.24	204.02	1815.70	213.32	2038.03	3.26	5.15
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Ca	300	990.75	4.03	337.28	421.60	231.88			32.806	10.04	2494.6	8.257	8.270	8.512	2043.07	122.5	7.67	4.86	1719.8	5.42	8	5.27	118.74	119.20	1709.00	323.33	2037.60	4.94	7.80
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Ca	450	664.01	2.36	63.24	368.90	231.88			32.800	10.05	2611.0	8.360	8.380	8.625	2054.60	90.9	9.57	6.06	1654.2	4.02	8	3.90	87.92	88.25	1641.22	402.88	2048.00	6.16	9.73
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Ca	600	796.82	2.91	263.50	421.60	111.72			32.841	10.09	2735.8	8.467	8.494	8.741	2055.94	65.7	11.84	7.50	1562.3	2.90	8	2.81	63.51	63.76	1548.45	497.76	2049.02	7.61	12.01
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Ca	0	2289.26	11.01	347.82	1138.31	803.14			32.797	11.75	2206.9	7.847	7.847	8.049	2029.95	392.2	3.11	1.98	1884.6	16.40	8	16.13	384.28	385.73	1880.74	130.98	2027.85	2.01	3.16
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Ca	150	1743.30	9.46	453.22	811.57	478.51			32.803	11.74	2344.2	8.063	8.063	8.271	2047.66	228.2	5.04	3.20	1829.3	9.54	8	9.38	223.48	224.33	1823.76	211.72	2044.86	3.25	5.11
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Ca	300	2133.27	9.61	537.53	980.21	615.53			32.765	11.76	2495.8	8.246	8.258	8.470	2053.01	136.8	7.56	4.80	1734.3	5.72	8	5.62	133.86	134.37	1726.89	316.95	2049.46	4.86	7.66
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Ca	450	2818.37	12.40	1085.61	1096.15	636.61			32.782	11.76	2617.0	8.351	8.370	8.585	2067.75	101.2	9.48	6.03	1670.8	4.23	8	4.15	98.82	99.20	1661.46	397.55	2063.16	6.10	9.60
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Ca	600	2089.01	11.04	537.53	1064.53	486.94			32.833	11.75	2736.0	8.453	8.479	8.696	2069.67	74.2	11.63	7.39	1584.6	3.10	8	3.04	72.54	72.81	1575.02	487.03	2065.10	7.47	11.76
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Ca	0	864.27	6.68	295.12	526.99	42.16			32.778	12.90	2202.9	7.877	7.877	8.061	2013.13	378.3	3.32	2.11	1860.6	15.25	8	15.06	372.22	373.61	1857.25	138.99	2011.31	2.14	3.36
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Ca	150	3233.64	20.44	1117.23	1770.70	345.71			32.736	12.91	2344.1	8.066	8.066	8.255	2046.35	237.7	5.08	3.24	1826.4	9.58	8	9.45	233.58	234.45	1821.44	212.94	2043.84	3.28	5.15
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Ca	300	1119.34	7.13	432.14	632.39	54.81			32.763	12.91	2491.4	8.255	8.268	8.460	2042.31	139.3	7.69	4.89	1718.5	5.61	8	5.54	136.98	137.49	1712.64	321.21	2039.39	4.94	7.76
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Ca	450	623.96	3.76	179.18	368.90	75.89			32.777	12.93	2608.1	8.355	8.375	8.569	2056.84	104.6	9.53	6.07	1658.0	4.21	8	4.15	102.73	103.11	1650.72	398.37	2053.25	6.13	9.63
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Ca	600	2502.17	17.18	621.85	1475.58	404.73			32.805	12.93	2738.1	8.467	8.494	8.691	2058.13	74.4	11.90	7.58	1562.3	3.00	8	2.95	73.03	73.30	1554.17	497.06	2054.18	7.65	12.01
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Ca	0	1521.96	9.13	885.35	632.39	4.22			32.798	14.72	2199.6	7.895	7.895	8.050	2001.75	387.3	3.46	2.21	1844.0	14.76	8	14.66	383.41	384.80	1841.61	144.23	2000.51	2.23	3.49
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Ca	150	857.95	5.54	379.44	474.29	4.22			32.762	14.72	2345.7	8.100	8.100	8.259	2027.43	232.6	5.44	3.48	1793.5	8.87	8	8.81	230.39	231.23	1790.39	226.69	2025.89	3.50	5.48
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Ca	300	600.77	3.13	368.90	221.34	10.54			32.787	14.71	2487.2	8.266	8.280	8.442	2029.67	144.7	7.86	5.02	1699.2	5.52	8	5.47	143.08	143.60	1694.87	327.12	2027.47	5.06	7.91
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Ca	450	463.76	2.39	168.64	284.58	10.54			32.786	14.74	2607.6	8.369	8.390	8.554	2043.86	107.8	9.78	6.25	1635.2	4.10	8	4.06	106.31	106.69	1629.36	407.46	2040.88	6.30	9.86
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Ca	600	685.09	5.32	221.34	453.22	10.54			32.812	14.76	2737.7	8.483	8.511	8.677	2042.08	76.0	12.21	7.81	1534.0	2.89	8	2.87	75.13	75.40	1528.38	508.05	2039.29	7.85	12.29
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Ca	0	690.06	5.34	388.92	301.14	0.00			32.806	15.38	2197.7	7.875	7.875	8.019	2009.02	420.0	3.33	2.13	1855.7	15.69	8	15.61	416.36	417.87	1853.71	138.67	2007.99	2.15	3.36
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Ca	150	684.59	7.36	246.64	431.63	6.32			32.750	15.38	2349.9	8.076	8.076	8.224	2045.81	256.8	5.23	3.35	1819.9	9.60	8	9.55	254.64	255.56	1817.24	217.66	2044.44	3.37	5.27
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Ca	300	81.81	0.43	18.97	50.19	12.65			32.769	15.38	2486.3	8.250	8.263	8.414	2041.83	156.8	7.65	4.90	1719.7	5.86	8	5.82	155.20	155.76	1715.49	318.38	2039.70	4.93	7.70
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Ca	450	569.51	2.92	199.20	351.33	18.97			32.784	15.38	2608.0	8.356	8.376	8.529	2055.93	115.8	9.60	6.14	1654.8	4.33	8	4.29	114.53	114.95	1649.94	399.20	2053.43	6.18	9.66
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Ca	600	525.35	4.72	123.33	391.48	10.54			32.806	15.38	2739.7	8.467	8.494	8.649	2059.55	82.8	11.96	7.66	1561.9	3.09	8	3.07	81.99	82.29	1557.23	496.94	2057.25	7.70	12.02
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Si	0	1254.25	3.44	143.34	834.76	276.15			32.760	9.63	2205.9	7.847	7.847	8.085	2027.62	358.9	3.08	1.95	1883.9	16.09	8	15.66	348.04	349.39	1879.42	130.75	2025.83	2.00	3.16
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Si	150	1658.98	4.48	303.55	1020.26	335.17			32.750	9.63	2346.4	8.088	8.089	8.334	2031.90	193.6	5.25	3.32	1805.8	8.68	8	8.45	187.84	188.57	1799.59	222.07	2030.11	3.39	5.36
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Si	300	1458.72	4.02	118.05	952.81	387.87			32.780	9.63	2481.8	8.250	8.263	8.512	2033.89	122.2	7.52	4.76	1716.9	5.48	8	5.33	118.36	118.82	1708.51	317.86	2031.70	4.86	7.67
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Si	450	1557.80	3.90	236.09	1028.69	293.01			32.790	9.62	2619.2	8.390	8.412	8.666	2029.00	80.5	10.03	6.35	1609.7	3.61	8	3.50	77.70	78.00	1599.08	424.06	2026.64	6.48	10.24
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	7	I	Si	600	1574.66	4.56	160.21	1155.17	259.28			32.820	9.62	2740.7	8.483	8.511	8.768	2038.21	60.6	12.10	7.66	1534.1	2.72	8	2.63	58.48	58.70	1522.31	510.94	2035.88	7.81	12.33
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Si	0	417.38	1.03	63.24	311.98	42.16			32.766	10.28	2197.7	7.841	7.841	8.067	2022.49	373.3	3.04	1.93	1880.2	16.38	8	16.03	364.00	365.40	1876.79	128.53	2021.35	1.97	3.10
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Si	150	1138.31	2.73	10.54	615.53	512.24			32.769	10.29	2339.9	8.082	8.082	8.316	2030.11	202.5	5.18	3.28	1806.7	8.88	8	8.67	196.88	197.64	1800.88	218.91	2028.46	3.35	5.29
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Si	300	874.81	2.11	63.24	371.00	440.57			32.793	10.30	2475.4	8.245	8.257	8.495	2032.21	127.5	7.45	4.72	1718.1	5.59	8	5.45	123.85	124.32	1710.70	314.32	2030.47	4.81	7.59
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Si	450	1176.25	2.95	52.70	438.46	685.09			32.800	10.31	2615.8	8.388	8.410	8.651	2027.88	83.4	10.00	6.34	1609.8	3.65	8	3.56	80.96	81.27	1601.52	421.68	2026.76	6.45	10.18
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	11	I	Si	600	1209.98	3.58	84.32	556.51	569.15			32.834	10.30	2739.0	8.484	8.513	8.756	2035.66	62.2	12.12	7.68	1530.6	2.72	8	2.66	60.40	60.63	1521.81	510.50	2034.97	7.81	12.32
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Si	0	173.38	0.46	63.24	65.87	44.27			32.794	10.13	2195.8	7.830	7.830	8.059	2025.27	381.9	2.97	1.88	1885.4	16.83	8	16.43	371.24	372.67	1881.44	125.84	2023.70	1.92	3.04
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Si	150	1369.66	2.94	347.82	935.42	86.43			32.783	10.10	2340.3	8.072	8.072	8.309	2036.32	206.8	5.08	3.22	1816.6	9.12	8	8.90	200.87	201.65	1810.65	215.04	2034.59	3.29	5.19
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Si	300	1917.73	4.24	400.52	1304.31	212.91			32.797	10.14	2477.6	8.234	8.245	8.486	2042.67	131.3	7.31	4.63	1734.1	5.79	8	5.64	127.44	127.94	1726.34	308.78	2040.76	4.72	7.45
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Si	450	664.01	1.67	84.32	368.90	210.80			32.799	10.14	2619.8	8.379	8.400	8.644	2039.54	85.5	9.88	6.26	1626.6	3.77	8	3.67	83.03	83.35	1618.19	416.45	2038.31	6.37	10.05
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	19	I	Si	600	820.53	2.30	189.72	487.47	143.34			32.828	10.15	2747.4	8.482	8.510	8.757	2044.37	62.4	12.12	7.68	1539.2	2.75	8	2.67	60.49	60.72	1529.34	511.11	2043.12	7.82	12.34
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Si	0	78.00	0.26	0.00	42.16	35.84			32.797	10.04	2196.3	7.820	7.820	8.050	2030.03	391.2	2.91	1.84	1892.2	17.29	8	16.87	380.14	381.61	1888.29	123.28	2028.44	1.89	2.98
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Si	150	944.37	2.93	210.80	484.83	248.74			32.788	10.03	2338.6	8.065	8.065	8.303	2038.89	210.2	5.01	3.18	1821.9	9.30	8	9.06	204.15	204.94	1815.93	212.15	2037.14	3.24	5.12
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Si	300	2105.87	6.26	389.98	990.75	725.14			32.806	10.05	2475.8	8.227	8.238	8.480	2046.53	133.8	7.21	4.57	1741.8	5.91	8	5.75	129.69	130.19	1733.86	304.85	2044.47	4.66	7.36
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Si	450	1250.03	4.39	263.50	685.09	301.44			32.812	10.06	2619.7	8.377	8.398	8.643	2041.17	85.8	9.84	6.24	1629.5	3.79	8	3.70	83.32	83.64	1621.22	415.10	2040.01	6.35	10.02
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	23	I	Si	600	1711.68	6.16	411.06	1053.99	246.63			32.843	10.07	2747.2	8.478	8.506	8.754	2048.09	63.1	12.05	7.64	1545.7	2.79	8	2.71	61.12	61.36	1535.70	508.29	2046.70	7.77	12.27
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Si	0	562.83	2.38	126.48	316.20	120.15			32.750	11.77	2194.8	7.818	7.818	8.019	2029.55	422.0	2.91	1.85	1891.2	17.64	8	17.36	413.67	415.24	1888.89	122.73	2028.98	1.88	2.96
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Si	150	2616.00	11.65	1011.83	1159.39	444.78			32.751	11.76	2338.3	8.053	8.053	8.261	2045.75	234.0	4.93	3.13	1831.9	9.78	8	9.63	229.35	230.22	1828.48	207.33	2045.44	3.18	5.01
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Si	300	1621.03	6.46	453.22	716.71	451.11			32.806	11.74	2477.2	8.221	8.232	8.444	2052.29	146.6	7.17	4.56	1749.2	6.13	8	6.03	143.51	144.05	1744.27	301.60	2051.89	4.63	7.28
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Si	450	3010.19	13.32	927.51	1443.96	638.72			32.800	11.75	2619.4	8.365	8.385	8.601	2051.81	96.2	9.69	6.16	1646.5	4.02	8	3.95	94.04	94.39	1640.59	407.31	2051.84	6.25	9.84
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	33	II	Si	600	1433.42	8.23	242.42	621.85	569.15			32.820	11.75	2749.1	8.467	8.494	8.712	2060.65	70.7	11.90	7.56	1564.6	2.96	8	2.90	69.19	69.45	1558.62	499.89	2061.41	7.67	12.07
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Si	0	8817.67	42.80	3436.00	4114.77	1266.89			32.751	12.90	2194.3	7.840	7.840	8.023	2019.71	416.3	3.06	1.95	1876.2	16.79	8	16.60	410.10	411.62	1874.61	128.45	2019.66	1.98	3.10
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Si	150	847.41	4.03	31.62	716.71	99.07			32.737	12.91	2332.5	8.105	8.108	8.297	2008.30	209.4	5.45	3.47	1774.4	8.44	8	8.35	206.29	207.06	1771.95	228.20	2008.50	3.51	5.52
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Si	300	419.49	2.25	21.08	305.66	92.75			32.774	12.92	2477.4	8.237	8.249	8.441	2040.52	146.5	7.40	4.71	1728.5	5.90	8	5.83	144.15	144.69	1725.16	309.91	2040.90	4.77	7.49
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Si	450	261.39	1.90	147.56	0.00	113.83			32.784	12.93	2618.6	8.383	8.405	8.600	2035.16	95.1	9.99	6.36	1618.0	3.83	8	3.78	93.52	93.87	1613.95	418.39	2036.12	6.44	10.11
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	41	II	Si	600	4525.83	29.94	1433.42	2371.47	720.93			32.815	12.92	2750.4	8.477	8.505	8.702	2052.02	71.8	12.10	7.71	1548.0	2.89	8	2.86	70.76	71.02	1544.65	506.61	2054.11	7.79	12.24
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Si	0	221.34	3.27	31.62	189.72	0.00			32.770	14.71	2196.0	7.888	7.888	8.043	1999.92	393.6	3.40	2.17	1844.2	15.01	8	14.93	390.08	391.50	1843.64	141.93	2000.50	2.19	3.43
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Si	150	406.84	2.04	0.00	389.98	16.86			32.765	14.71	2329.5	8.100	8.100	8.260	2010.05	230.4	5.40	3.45	1778.0	8.79	8	8.73	228.08	228.90	1776.61	225.39	2010.73	3.48	5.45
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Si	300	168.64	0.99	0.00	158.10	10.54			32.790	14.72	2472.9	8.264	8.277	8.440	2015.24	144.5	7.77	4.97	1688.4	5.51	8	5.47	143.06	143.59	1686.78	324.21	2016.46	5.01	7.84
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Si	450	3599.37	10.19	516.45	3082.92	0.00			32.795	14.76	2621.5	8.403	8.426	8.590	2019.25	96.1	10.35	6.61	1587.7	3.66	8	3.64	95.29	95.64	1586.60	431.48	2021.72	6.67	10.44
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	49	II	Si	600	75.89	1.24	52.70	21.08	2.11			32.822	14.77	2751.2	8.521	8.552	8.719	2009.09	66.4	12.91	8.25	1472.8	2.53	8	2.51	65.60	65.84	1470.11	538.63	2011.24	8.33	13.02
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Si	0	498.59	6.72	37.95	441.67	18.97			32.761	15.39	2197.4	7.893	7.893	8.038	1999.15	399.5	3.45	2.21	1841.8	14.92	8	14.84	395.87	397.29	1840.83	143.77	1999.44	2.23	3.48
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Si	150	426.77	2.63	9.49	381.44	35.84			32.749	15.37	2332.0	8.087	8.087	8.236	2020.47	246.4	5.29	3.39	1792.6	9.21	8	9.16	244.19	245.07	1791.58	220.51	2021.25	3.42	5.34
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Si	300	976.00	8.52	18.97	927.51	29.51			32.762	15.38	2469.9	8.249	8.261	8.413	2025.14	156.0	7.57	4.85	1706.2	5.83	8	5.80	154.59	155.15	1704.85	315.63	2026.28	4.89	7.64
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Si	450	4568.46	11.56	531.21	2471.02	1566.23			32.791	15.37	2622.2	8.380	8.401	8.555	2041.42	107.0	10.01	6.40	1623.8	4.00	8	3.98	106.10	106.48	1622.88	416.79	2043.64	6.45	10.08
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	53	II	Si	600	545.16	6.87	18.97	521.98	4.22			32.821	15.37	2749.9	8.508	8.538	8.694	2021.58	71.5	12.69	8.12	1494.3	2.67	8	2.66	70.87	71.13	1493.08	528.47	2024.22	8.18	12.78
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	0						6
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	0						7
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	0						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	0						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	150						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	150						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	150						6
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	150						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	300						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	300						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	300						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	300						2
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	450						3
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	450						3
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	450						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	450						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	600						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	600						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	600						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	600						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	0						6
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	0						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	0						3
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	0						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	150						6
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	150						6
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	150						6
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	150						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	300						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	300						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	300						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	300						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	450						4
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	450						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	450						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	450						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	600						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	600						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	600						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	600						5
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	0							6.41
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	150							3.82
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	300							5.19
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	450							4.30
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Ca	600							3.59
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	0							4.82
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	150							7.73
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	300							2.34
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	450							4.44
laboratory	Oikopleura dioica	Oikopleura dioica	Oikopleura dioica	36		Si	600							5.11