Citation:

PANGAEA.992011

Name: GDGTs of soil and lacustrine sediments in Cameroon
Published: 2026-06-29
License: https://creativecommons.org/licenses/by/4.0/
Keywords: Cameroon; GDGT; Holocene; lake Mbalang; modern sediments

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Published: 2026-06-29 • DOI registered: 2026-07-28

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Abstract:

This file contains the biomarker data from the paper Ménot et al., 2025 Organic Geochemistry (https://doi.org/10.1016/j.orggeochem.2025.104982). African low-latitude regions tend to be underrepresented in global continental temperature reconstructions, limiting both our understanding and the reliability of predictions of past and future changes in temperature and precipitation in those parts of the world. The lack of continuous sedimentary archives and quantitative temperature proxies further complicates this issue, especially outside Eastern Africa. Here, we use data collected in Cameroon lakes to assess branched glycerol dialkyl glycerol tetraether (brGDGT)-based temperature proxies and examine potential confounding variables impacting temperature reconstructions. By analyzing GDGT distributions in soil watershed, water column, and surface sediment samples from a total of 11 Cameroon lakes, we could verify that the degree of methylation of brGDGTs (through the MBT′5Me) can serve as a reliable indicator for reconstructing mean annual air temperatures. Additionally, we could confirm that surface water conductivity represents a controlling factor for the brGDGT assemblage in surface sediments of crater lakes. Moreover, we provide the first reconstruction of Mid- to Late-Holocene GDGT-based air temperatures for a crater lake in Cameroon (Central Africa), revealing a temperature decrease of 2.5 °C over the last 7000 years, which agrees with recently published records for East Africa but exceeds current model predictions. These discrepancies highlight the need for additional studies to focus on this geographically underrepresented area.

Keyword(s):

Cameroon; GDGT; Holocene; lake Mbalang; modern sediments

Related to:

Ménot, Guillemette; Ansanay-Alex, Salomé; Schwab, Valérie F; Todou, Gilbert; Séné, Olivier; Onana, Jean-Michel; Gleixner, Gerd; Sachse, Dirk; Garcin, Yannick (2025): Mid- to Late-Holocene branched GDGT-based air temperatures from a crater lake in Cameroon (Central Africa). Organic Geochemistry, 204, 104982, https://doi.org/10.1016/j.orggeochem.2025.104982

Funding:

Agence Nationale de la Recherche (ANR), grant/award no. ANR-18-CE01-0005: ANR TAPIOCA

German Research Foundation (DFG), grant/award no. 160487544

Coverage:

Median Latitude: 6.734127 * Median Longitude: 11.848582 * South-bound Latitude: 3.768600 * West-bound Longitude: 8.980500 * North-bound Latitude: 11.054700 * East-bound Longitude: 14.145700

Minimum Elevation: 17.0 m * Maximum Elevation: 1911.0 m

Event(s):

ASSO * Latitude: 6.624500 * Longitude: 12.981600 * Elevation: 896.0 m * Comment: Assom lake basin

BALE * Latitude: 5.550300 * Longitude: 10.421100 * Elevation: 1376.0 m * Comment: Baleng lake basin

BARO * Latitude: 4.654400 * Longitude: 9.408600 * Elevation: 304.0 m * Comment: Barombi Mbo lake basin

Parameter(s):

#	Name	Short Name	Unit	Principal Investigator	Method/Device	Comment
1	Event label	Event		Ménot, Guillemette
2	DEPTH, sediment/rock	Depth sed	m	Ménot, Guillemette	see reference(s)	Geocode
3	Depth, top/min	Depth top	m	Ménot, Guillemette	see reference(s)
4	Depth, bottom/max	Depth bot	m	Ménot, Guillemette	see reference(s)
5	DEPTH, water	Depth water	m	Ménot, Guillemette	see reference(s)	Geocode
6	DEPTH, soil	Depth soil	m	Ménot, Guillemette	see reference(s)	Geocode
7	Depth, soil, minimum	Depth soil min	m	Ménot, Guillemette	see reference(s)
8	Depth, soil, maximum	Depth soil max	m	Ménot, Guillemette	see reference(s)
9	Sample type	Samp type		Ménot, Guillemette	see reference(s)
10	Lake	Lake		Ménot, Guillemette	see reference(s)	basin
11	Branched glycerol dialkyl glycerol tetraether (peak area)	brGDGT peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200	sum
12	Isoprenoid glycerol dialkyl glycerol tetraether (peak area)	iso GDGT peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200	sum
13	Crenarchaeol (peak area)	Cren peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
14	Branched glycerol dialkyl glycerol tetraether, IIIa (peak area)	brGDGT IIIa peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
15	Branched glycerol dialkyl glycerol tetraether, IIIa' (peak area)	brGDGT IIIa' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
16	Branched glycerol dialkyl glycerol tetraether, IIIa'' (peak area)	brGDGT IIIa'' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
17	Branched glycerol dialkyl glycerol tetraether, IIIb (peak area)	brGDGT IIIb peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
18	Branched glycerol dialkyl glycerol tetraether, IIIb' (peak area)	brGDGT IIIb' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
19	Branched glycerol dialkyl glycerol tetraether, IIIb'' (peak area)	brGDGT IIIb'' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
20	Branched glycerol dialkyl glycerol tetraether, IIIc (peak area)	brGDGT IIIc peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
21	Branched glycerol dialkyl glycerol tetraether, IIIc' (peak area)	brGDGT IIIc' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
22	Branched glycerol dialkyl glycerol tetraether, IIa (peak area)	brGDGT IIa peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
23	Branched glycerol dialkyl glycerol tetraether, IIa' (peak area)	brGDGT IIa' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
24	Branched glycerol dialkyl glycerol tetraether, IIb (peak area)	brGDGT IIb peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
25	Branched glycerol dialkyl glycerol tetraether, IIb' (peak area)	brGDGT IIb' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
26	Branched glycerol dialkyl glycerol tetraether, IIc (peak area)	brGDGT IIc peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
27	Branched glycerol dialkyl glycerol tetraether, IIc' (peak area)	brGDGT IIc' peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
28	Branched glycerol dialkyl glycerol tetraether, Ia (peak area)	brGDGT Ia peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
29	Branched glycerol dialkyl glycerol tetraether, Ib (peak area)	brGDGT Ib peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
30	Branched glycerol dialkyl glycerol tetraether, Ic (peak area)	brGDGT Ic peak area		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
31	Isoprenoid acyclic glycerol dialkyl glycerol tetraether, fractional abundance	iso GDGT-0		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200	total
32	Branched glycerol dialkyl glycerol tetraether, fractional abundance	brGDGT		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200	total
33	Branched glycerol dialkyl glycerol tetraether, IIIa, fractional abundance	brGDGT IIIa		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
34	Branched glycerol dialkyl glycerol tetraether, IIIa', fractional abundance	brGDGT IIIa'		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
35	Branched glycerol dialkyl glycerol tetraether, IIIa'', fractional abundance	brGDGT IIIa''		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
36	Branched glycerol dialkyl glycerol tetraether, IIIb, fractional abundance	brGDGT IIIb		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
37	Branched glycerol dialkyl glycerol tetraether, IIIb', fractional abundance	brGDGT IIIb'		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
38	Branched glycerol dialkyl glycerol tetraether, IIIb'', fractional abundance	brGDGT IIIb''		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
39	Branched glycerol dialkyl glycerol tetraether, IIIc, fractional abundance	brGDGT IIIc		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
40	Branched glycerol dialkyl glycerol tetraether, IIIc', fractional abundance	brGDGT IIIc'		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
41	Branched glycerol dialkyl glycerol tetraether, IIa, fractional abundance	brGDGT IIa		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
42	Branched glycerol dialkyl glycerol tetraether, IIa', fractional abundance	brGDGT IIa'		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
43	Branched glycerol dialkyl glycerol tetraether, IIb, fractional abundance	brGDGT IIb		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
44	Branched glycerol dialkyl glycerol tetraether, IIb', fractional abundance	brGDGT IIb'		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
45	Branched glycerol dialkyl glycerol tetraether, IIc, fractional abundance	brGDGT IIc		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
46	Branched glycerol dialkyl glycerol tetraether, IIc', fractional abundance	brGDGT IIc'		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
47	Branched glycerol dialkyl glycerol tetraether, Ia, fractional abundance	brGDGT Ia		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
48	Branched glycerol dialkyl glycerol tetraether, Ib, fractional abundance	brGDGT Ib		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
49	Branched glycerol dialkyl glycerol tetraether, Ic, fractional abundance	brGDGT Ic		Ménot, Guillemette	High Performance Liquid Chromatography Mass Spectrometry (HPLC-APCI-MS), Agilent 1200
50	Branched and isoprenoid tetraether index	BIT		Ménot, Guillemette	see reference(s)
51	Methylation index of 5-methyl branched tetraethers	MBT'5ME		Ménot, Guillemette	see reference(s)
52	Isomer ratio of 6-methyl branched glycerol dialkyl glycerol tetraethers	IR6me		Ménot, Guillemette	see reference(s)
53	Cyclization ratio of branched tetraethers prime	CBT'		Ménot, Guillemette	see reference(s)
54	Cyclization ratio of branched tetraethers of 5-methyl branched glycerol dialkyl glycerol tetraether	CBT5ME		Ménot, Guillemette	see reference(s)

License:

Creative Commons Attribution 4.0 International (CC-BY-4.0)

Status:

Curation Level: Enhanced curation (CurationLevelC)

Size:

1514 data points

Data

Download dataset as tab-delimited text — use the following character encoding:

1 Event	2 Depth sed [m]	3 Depth top [m]	4 Depth bot [m]	5 Depth water [m]	6 Depth soil [m]	7 Depth soil min [m]	8 Depth soil max [m]	9 Samp type	10 Lake	11 brGDGT peak area	12 iso GDGT peak area	13 Cren peak area	14 brGDGT IIIa peak area	15 brGDGT IIIa' peak area	16 brGDGT IIIa'' peak area	17 brGDGT IIIb peak area	18 brGDGT IIIb' peak area	19 brGDGT IIIb'' peak area	20 brGDGT IIIc peak area	21 brGDGT IIIc' peak area	22 brGDGT IIa peak area	23 brGDGT IIa' peak area	24 brGDGT IIb peak area	25 brGDGT IIb' peak area	26 brGDGT IIc peak area	27 brGDGT IIc' peak area	28 brGDGT Ia peak area	29 brGDGT Ib peak area	30 brGDGT Ic peak area	31 iso GDGT-0	32 brGDGT	33 brGDGT IIIa	34 brGDGT IIIa'	35 brGDGT IIIa''	36 brGDGT IIIb	38 brGDGT IIIb''	39 brGDGT IIIc	40 brGDGT IIIc'	41 brGDGT IIa	42 brGDGT IIa'	43 brGDGT IIb	44 brGDGT IIb'	45 brGDGT IIc	46 brGDGT IIc'	47 brGDGT Ia	48 brGDGT Ib	49 brGDGT Ic	50 BIT	51 MBT'5ME	52 IR6me	53 CBT'	54 CBT5ME
OSSA	0.025	0.01	0.04					Sediment	Ossa	6332085	88173	13451	64735	33310	186049	1236	2226	0	3144	0	724447	310534	66549	47514	11036	2344	4152042	555172	83572	1.39	98.61	1.02	0.53	2.94	0.02	0.00	0.05	0.00	11.44	4.90	1.05	0.75	0.17	0.04	65.57	8.77	1.32	1.00	0.85	0.31	-1.02	-0.89
DEBU	0.025	0.01	0.04					Sediment	Debundscha	297714780	32598565	6930792	1503361	1510751	5201505	175078	274155	278072	75409	0	16039586	11798509	747113	3322625	163651	102927	207734048	13833884	2355541	10.95	89.05	0.50	0.51	1.75	0.06	0.09	0.03	0.00	5.39	3.96	0.25	1.12	0.05	0.03	69.78	4.65	0.79	0.97	0.92	0.47	-1.07	-1.19
BARO	0.025	0.01	0.04					Sediment	Barombi Mbo	123323747	44610451	17433442	1080700	856819	1575313	68853	115589	70093	42362	0	6661750	6838115	890120	2012858	100394	136702	48909316	7870601	1483713	36.17	63.83	0.88	0.69	1.28	0.06	0.06	0.03	0.00	5.40	5.54	0.72	1.63	0.08	0.11	39.66	6.38	1.20	0.82	0.87	0.53	-0.70	-0.80
MANE	0.025	0.01	0.04					Sediment	Manengouba	85223904	46054973	14430102	1211979	875593	910596	70919	81022	68013	20114	9073	7916573	3820015	1149425	652942	239067	69895	18677110	2595983	800615	54.04	45.96	1.42	1.03	1.07	0.08	0.08	0.02	0.01	9.29	4.48	1.35	0.77	0.28	0.08	21.92	3.05	0.94	0.73	0.68	0.34	-0.65	-0.85
BALE	0.025	0.01	0.04					Sediment	Baleng	495611257	105978457	3767867	6326855	20566312	5370977	662096	3380458	668237	103832	627807	35208796	83287104	11835792	46902448	710278	2749311	118346088	45526808	7359602	21.38	78.62	1.28	4.15	1.08	0.13	0.13	0.02	0.13	7.10	16.80	2.39	9.46	0.14	0.55	23.88	9.19	1.48	0.99	0.71	0.74	0.00	-0.43
ASSO	0.025	0.01	0.04					Sediment	Assom	39838170	3345700	18400	344686	457906	1253174	101601	55509	38789	49508	0	4182020	2603645	577198	340852	120529	8654	22943268	2807353	607779	8.40	91.60	0.87	1.15	3.15	0.26	0.10	0.12	0.00	10.50	6.54	1.45	0.86	0.30	0.02	57.59	7.05	1.53	1.00	0.83	0.39	-0.83	-0.90
TABE	0.025	0.01	0.04					Sediment	Tabéré	2047778	500651	6696	57334	66003	31217	0	0	0	3180	0	203735	206812	37829	68954	11563	2982	688480	133776	35262	24.45	75.55	2.80	3.22	1.52	0.00	0.00	0.16	0.00	9.95	10.10	1.85	3.37	0.56	0.15	33.62	6.53	1.72	1.00	0.73	0.53	-0.40	-0.72
TIZO	0.025	0.01	0.04					Sediment	Tizong	320251962	85484704	9862933	7549729	14708472	2315414	646495	756865	332405	56822	90163	27112026	60494096	10790469	10689420	330126	564230	75214784	20433496	2682245	26.69	73.31	2.36	4.59	0.72	0.20	0.10	0.02	0.03	8.47	18.89	3.37	3.34	0.10	0.18	23.49	6.38	0.84	0.96	0.65	0.65	-0.09	-0.52
MAMG	0.025	0.01	0.04					Sediment	Mamguiewa	126370428	12987867	592637	993795	2379875	1774140	205931	494736	152435	180853	118156	9031698	14277952	4415006	9626403	331354	1410530	42482760	22050252	3456685	10.28	89.72	0.79	1.88	1.40	0.16	0.12	0.14	0.09	7.15	11.30	3.49	7.62	0.26	1.12	33.62	17.45	2.74	0.99	0.81	0.65	-0.23	-0.29
RHUM	0.025	0.01	0.04					Sediment	Rhumsiki	88425714	23851924	1842419	1244754	1703319	619073	36038	38816	13982	7817	8028	10854818	9059393	922592	724825	33118	16586	35687552	3304230	298852	26.97	73.03	1.41	1.93	0.70	0.04	0.02	0.01	0.01	12.28	10.25	1.04	0.82	0.04	0.02	40.36	3.74	0.34	0.97	0.74	0.47	-0.61	-1.04
MORA	0.025	0.01	0.04					Sediment	Mora	25041548	2766765	451939	200315	307463	59951	0	0	0	1583	0	3651039	2325834	39612	20017	0	0	15482989	172592	13388	11.05	88.95	0.80	1.23	0.24	0.00	0.00	0.01	0.00	14.58	9.29	0.16	0.08	0.00	0.00	61.83	0.69	0.05	0.98	0.80	0.41	-0.86	-1.96
OSSA				1				Filter	Ossa	7803174	632450	73717	37025	52324	39354	100938	36630	0	98086	0	482628	329140	19488	37886	0	0	5679637	195115	62474	8.11	91.89	0.47	0.67	0.50	1.29	0.00	1.26	0.00	6.19	4.22	0.25	0.49	0.00	0.00	72.79	2.50	0.80	0.99	0.91	0.36	-1.11	-1.46
DEBU				1				Filter	Debundscha	1701484	151470	32593	3119	11826	5218	1990	0	0	0	0	41586	129059	2337	16484	3842	1494	1263610	59510	9938	8.90	91.10	0.18	0.70	0.31	0.12	0.00	0.00	0.00	2.44	7.59	0.14	0.97	0.23	0.09	74.27	3.50	0.58	0.98	0.96	0.76	-0.89	-1.32
BARO				1				Filter	Barombi Mbo	2644186	480845	157484	10715	29389	16401	1634	4163	0	0	0	110207	248983	16608	89552	3462	6818	1353216	230691	41502	18.18	81.82	0.41	1.11	0.62	0.06	0.00	0.00	0.00	4.17	9.42	0.63	3.39	0.13	0.26	51.18	8.72	1.57	0.93	0.91	0.73	-0.55	-0.77
MANE				1				Filter	Manengouba	15589701	6568552	2704663	203261	209559	95877	12482	0	0	0	0	1748083	1261896	112824	192534	16679	14383	4626985	431013	95573	42.13	57.87	1.30	1.34	0.62	0.08	0.00	0.00	0.00	11.21	8.09	0.72	1.24	0.11	0.09	29.68	2.76	0.61	0.77	0.71	0.45	-0.57	-1.07
BALE				1				Filter	Baleng	530608	101335	9430	44824	0	0	4046	0	0	0	0	29466	99617	5157	41488	0	0	151811	38323	14543	19.10	80.90	8.45	0.00	0.00	0.76	0.00	0.00	0.00	5.55	18.77	0.97	7.82	0.00	0.00	28.61	7.22	2.74	0.98	0.86	0.63	-0.16	-0.62
ASSO				1				Filter	Assom	15121369	1942102	156672	104577	180252	175729	5798	15636	9417	0	0	1310342	1183738	81920	280582	14870	8795	9127015	594115	86482	12.84	87.16	0.69	1.19	1.16	0.04	0.06	0.00	0.00	8.67	7.83	0.54	1.86	0.10	0.06	60.36	3.93	0.57	0.99	0.86	0.52	-0.78	-1.19
TABE				1				Filter	Tabéré	9979896	1532440	26712	133626	345401	78438	6119	19698	7641	0	0	1138033	1793973	73431	418582	8449	20150	3814939	525661	63317	15.36	84.64	1.34	3.46	0.79	0.06	0.08	0.00	0.00	11.40	17.98	0.74	4.19	0.08	0.20	38.23	5.27	0.63	1.00	0.74	0.66	-0.28	-0.92
TIZO				1				Filter	Tizong	2645819	530817	20884	21257	153479	19898	3429	9631	4778	3263	4191	105481	675240	55629	176311	5747	13197	627928	200301	35241	20.06	79.94	0.80	5.80	0.75	0.13	0.18	0.12	0.16	3.99	25.52	2.10	6.66	0.22	0.50	23.73	7.57	1.33	0.99	0.73	0.84	0.15	-0.46
MAMG				1				Filter	Mamguiewa	6558568	642765	149708	34244	174336	34447	2724	27386	4893	4945	7471	313719	1381437	51825	591336	4235	49211	2344615	809747	79231	9.80	90.20	0.52	2.66	0.53	0.04	0.07	0.08	0.11	4.78	21.06	0.79	9.02	0.06	0.75	35.75	12.35	1.21	0.98	0.86	0.84	-0.07	-0.49
RHUM				1				Filter	Rhumsiki	10753669	3033613	217869	96423	204057	26719	8384	2255	7657	0	0	1165960	1131352	55126	134985	0	0	4515864	347270	24007	28.21	71.79	0.90	1.90	0.25	0.08	0.07	0.00	0.00	10.84	10.52	0.51	1.26	0.00	0.00	41.99	3.23	0.22	0.97	0.77	0.53	-0.59	-1.15
MORA				1				Filter	Mora	55082875	14966500	538606	404342	1736210	85252	0	0	0	0	0	5430885	9056383	58486	125090	0	0	22881406	338322	0	27.17	72.83	0.73	3.15	0.15	0.00	0.00	0.00	0.00	9.86	16.44	0.11	0.23	0.00	0.00	41.54	0.61	0.00	0.99	0.76	0.65	-0.42	-1.85
OSSA					0.025	0	0.05	Soil	Ossa	74773829	4245956	1324018	46917	11517	0	0	0	0	51073	3904	1484196	150275	28564	24002	40713	0	67229608	882227	574875	5.68	94.32	0.06	0.02	0.00	0.00	0.00	0.07	0.01	1.98	0.20	0.04	0.03	0.05	0.00	89.91	1.18	0.77	0.98	0.98	0.11	-1.95	-1.88
DEBU					0.025	0	0.05	Soil	Debundscha	272095294	9115749	3889773	539900	1091610	75879	0	0	0	0	0	10539724	13901206	2584092	2198309	140074	111767	181373104	40447696	9976185	3.35	96.65	0.20	0.40	0.03	0.00	0.00	0.00	0.00	3.87	5.11	0.95	0.81	0.05	0.04	66.66	14.87	3.67	0.99	0.94	0.56	-0.85	-0.65
BARO					0.025	0	0.05	Soil	Barombi Mbo	361996834	28910309	20627508	1150586	2222245	104559	279415	346577	131759	48357	41245	13157393	19783352	6020109	6194104	536511	360842	178461152	76691128	27557192	7.99	92.01	0.32	0.61	0.03	0.08	0.04	0.01	0.01	3.63	5.47	1.66	1.71	0.15	0.10	49.30	21.19	7.61	0.94	0.93	0.58	-0.54	-0.36
MANE					0.025	0	0.05	Soil	Manengouba	79983445	1738358	260164	654668	788468	0	104640	0	0	16838	0	10240149	6213144	184927	623702	68384	0	55566512	3337486	446169	2.17	97.83	0.82	0.99	0.00	0.13	0.00	0.02	0.00	12.80	7.77	0.23	0.78	0.09	0.00	69.47	4.17	0.56	1.00	0.84	0.41	-0.92	-1.27
ASSO					0.025	0	0.05	Soil	Assom	94446332	2723037	181833	47500	0	0	0	0	0	12891	0	1497379	275895	0	0	96364	0	87578200	1742172	472895	2.88	97.12	0.05	0.00	0.00	0.00	0.00	0.01	0.00	1.59	0.29	0.00	0.00	0.10	0.00	92.73	1.84	0.50	1.00	0.98	0.15	-2.08	-1.71
TABE					0.025	0	0.05	Soil	Tabéré	67690904	8723905	3436212	707829	1334325	77764	63995	134631	33335	33444	27521	2862464	12603179	887233	2948018	96627	148689	26131994	9133074	1742877	12.89	87.11	1.05	1.97	0.11	0.09	0.05	0.05	0.04	4.23	18.62	1.31	4.36	0.14	0.22	38.60	13.49	2.57	0.94	0.88	0.79	-0.20	-0.46
TIZO					0.025	0	0.05	Soil	Tizong	27623426	856698	128945	38073	179371	12192	0	0	0	14623	0	865317	2607727	115899	247946	27268	0	20060536	2250803	346972	3.10	96.90	0.14	0.65	0.04	0.00	0.00	0.05	0.00	3.13	9.44	0.42	0.90	0.10	0.00	72.62	8.15	1.26	1.00	0.95	0.75	-0.79	-0.95
MAMG					0.025	0	0.05	Soil	Mamguiewa	13467816	2811036	1031722	42934	63790	0	0	0	0	9535	0	476789	1068169	120855	39534	0	0	6921696	1719132	194347	20.87	79.13	0.32	0.47	0.00	0.00	0.00	0.07	0.00	3.54	7.93	0.90	0.29	0.00	0.00	51.39	12.76	1.44	0.91	0.93	0.64	-0.74	-0.60
RHUM					0.025	0	0.05	Soil	Rhumsiki	159710138	4553645	409953	442413	1233030	419658	18527	16783	0	46711	0	31669842	13706449	519345	516971	16817	0	104098712	2231903	219331	2.85	97.15	0.28	0.77	0.26	0.01	0.00	0.03	0.00	19.83	8.58	0.33	0.32	0.01	0.00	65.18	1.40	0.14	1.00	0.76	0.32	-0.94	-1.69
MORA					0.025	0	0.05	Soil	Mora	87878303	5442737	2515715	99913	203333	6365	0	0	0	42636	0	3774758	2629552	17029	10149	15767	0	75180464	394490	61111	6.19	93.81	0.11	0.23	0.01	0.00	0.00	0.05	0.00	4.30	2.99	0.02	0.01	0.02	0.00	85.55	0.45	0.07	0.97	0.95	0.42	-1.43	-2.28