Citation:

PANGAEA.988290

Name: Microbial Nitrogen Cycling Genes in Arctic Streams
Published: 2026-02-09
License: https://creativecommons.org/licenses/by-sa/4.0/
Keywords: Arctic; diversity; nitrogen genes; Sediment; stream; water chemistry; Watershed

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Published: 2026-02-09 • DOI registered: 2026-02-09

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Abstract:

This study integrates molecular, environmental, and spatiotemporal data to examine the structure and function of streambed microbial communities across Arctic regions experiencing rapid climate change. Sediment samples were collected from 28 streams spanning four Arctic regions during summer 2021, including 15 Greenlandic streams that were sampled three times throughout the open-water season to capture temporal variation. Microbial community composition was characterized using 16S rRNA gene sequencing, while functional potential for nitrogen cycling was quantified through qPCR targeting six key genes involved in N-fixation (nifH), nitrification (amoA, nxrB), and denitrification (nirS, norB, nosZ). Environmental data included catchment characteristics, streamwater and sediment caractheristics. These molecular and environmental datasets were integrated with spatial and temporal sampling metadata to assess biogeographic and seasonal trends. Together, these data provide a comprehensive framework for understanding how Arctic stream microbial communities and nitrogen cycling functions respond to spatial heterogeneity and seasonal dynamics under ongoing climate change.

Keyword(s):

Arctic; diversity; nitrogen genes; Sediment; stream; water chemistry; Watershed

Supplement to:

Holmboe, Cecilie Marie Hartvig; Riis, Tenna; Han, Xingguo; Frossard, Aline; Romaní, Anna M; Kjær, Jakob B; Tank, Jennifer; Giménez-Grau, Pau; Hille, Erika; D'Acqui, Luigi P; Catalán, Núria; Pastor, Ada (accepted): Spatial and Temporal Variability of Microbial Nitrogen Cycling Genes in Arctic Streams. Global Biogeochemical Cycles

Related to:

Holmboe, Cecilie Marie Hartvig; Pastor, Ada; Riis, Tenna (2024): Stream Nitrogen Concentrations Across Arctic Vegetation Gradients. Global Biogeochemical Cycles, 38(2), e2023GB007840, https://doi.org/10.1029/2023GB007840

Funding:

Aarhus University (AU), grant/award no. AUFF-E-2020-9-16: Deciphering the role of rocks as nitrogen source for biological cycling in Arctic freshwaters (RockNrole)

Danish National Research Foundation (DG), grant/award no. DNRF185 : Center for Ice-Free Arctic Research (CIFAR)

Horizon 2020 (H2020), grant/award no. 730938: Nitrogen sources for Arctic freshwaters (NitroFresh)

Horizon 2020 (H2020), grant/award no. 839709 : Linking Chemical diversity and Reactivity of Arctic dissolved Organic Matter for its integration in Earth system model

Spanish Ministry of Science, Innovation and Universities (MICINN), grant/award no. RYC2022-036661-I

Villum Fonden, grant/award no. 00028351 : Nitrogen in rocks: an overlooked nutrient source in Arctic aquatic ecosystems

Coverage:

Median Latitude: 73.527082 * Median Longitude: -49.890874 * South-bound Latitude: 67.753333 * West-bound Longitude: -149.592708 * North-bound Latitude: 78.936310 * East-bound Longitude: 12.007480

Date/Time Start: 2021-07-09T00:00:00 * Date/Time End: 2021-09-17T00:00:00

Minimum Elevation: 5.7 m * Maximum Elevation: 890.4 m

Event(s):

Aucella_microbes * Latitude: 74.503270 * Longitude: -20.541790 * Date/Time Start: 2021-07-18T00:00:00 * Date/Time End: 2021-09-12T00:00:00 * Elevation: 153.9 m * Location: Northeast Greenland * Method/Device: Multiple investigations (MULT)

Austre-Bayelva_microbes * Latitude: 78.919940 * Longitude: 11.826250 * Date/Time: 2021-08-23T00:00:00 * Elevation: 52.5 m * Location: Svalbard * Method/Device: Multiple investigations (MULT)

Bayelva_microbes * Latitude: 78.920320 * Longitude: 11.795890 * Date/Time: 2021-08-27T00:00:00 * Elevation: 50.4 m * Location: Svalbard * Method/Device: Multiple investigations (MULT)

Parameter(s):

#	Name	Short Name	Unit	Principal Investigator	Comment
1	Event label	Event		Pastor, Ada
2	Sample code/label	Sample label		Pastor, Ada
3	Latitude of event	Latitude		Pastor, Ada
4	Longitude of event	Longitude		Pastor, Ada
5	DATE/TIME	Date/Time		Pastor, Ada	Geocode
6	Elevation of event	Elevation	m	Pastor, Ada
7	pH	pH		Pastor, Ada	streamwater
8	Temperature, water	Temp	°C	Pastor, Ada	streamwater
9	Conductivity, electrical	EC	µS/cm	Pastor, Ada	streamwater
10	Ammonium	[NH4]+	mg/l	Pastor, Ada	streamwater
11	Nitrate	[NO3]-	mg/l	Pastor, Ada	streamwater
12	Carbon, organic, dissolved	DOC	mg/l	Pastor, Ada	streamwater
13	Nitrogen, organic, dissolved	DON	mg/l	Pastor, Ada	streamwater
14	Nitrogen, total dissolved	TDN	mg/l	Pastor, Ada	streamwater
15	pH	pH		Pastor, Ada	sediment
16	Conductivity, electrical	EC	µS/cm	Pastor, Ada	sediment
17	Size fraction > 2 mm, gravel	>2 mm	%	Pastor, Ada	Sediment size >2000 m
18	Size fraction 2.000-0.630 mm, coarse sand	2000-630 µm	%	Pastor, Ada	Sediment size 2000-630 m
19	Size fraction 0.630-0.200 mm, medium sand	630-200 µm	%	Pastor, Ada	Sediment size 630-200 m
20	Size fraction 0.200-0.063 mm, fine sand	200-63 µm fS	%	Pastor, Ada	Sediment size 200 63 m
21	Loss on ignition	LOI	%	Pastor, Ada
22	Ammonium	[NH4]+	mg/l	Pastor, Ada	sediment
23	Nitrate	[NO3]-	mg/l	Pastor, Ada	sediment
24	Nitrogen	N	%	Pastor, Ada	sediment
25	Carbon	C	%	Pastor, Ada	sediment
26	Normalized Difference Vegetation Index	NDVI		Pastor, Ada
27	Catchment area	Catch area	km²	Pastor, Ada
28	Slope	Slope	deg	Pastor, Ada
29	16S rRNA gene, copy number, per unit sediment dry mass	16S rRNA gene/sed dm	#/g	Pastor, Ada
30	AOA-amoA gene, copy number, per unit sediment dry mass	AOA-amoA/sed dm	#/g	Pastor, Ada	Archaeal amoA gene (encoding ammonium monooxygenase) in sediment samples, quantified from extracted DNA
31	AOB-amoA gene, copy number, per unit sediment dry mass	AOB-amoA/sed dm	#/g	Pastor, Ada	Bacterial amoA gene (encoding ammonium monooxygenase) in sediment samples, quantified from extracted DNA
32	nifH gene, copy number, per unit sediment dry mass	nifH/sed dm	#/g	Pastor, Ada	nifH gene (encoding the nitrogenase enzyme) in sediment samples, quantified from extracted DNA
33	nrxB gene, copy number, per unit sediment dry mass	nrxB/sed dm	#/g	Pastor, Ada	nrxB gene (encoding the beta subunit of nitrite oxidoreductase) in sediment samples, quantified from extracted DNA
34	nosZ gene, copy number, per unit sediment dry mass	nosZ/sed dm	#/g	Pastor, Ada	nosZ gene (encoding nitrous oxide reductase) in sediment samples, quantified from extracted DNA
35	nirS gene, copy number, per unit sediment dry mass	nirS/sed dm	#/g	Pastor, Ada	nirS gene (encoding cytochrome cd1 nitrite reductase) in sediment samples, quantified from extracted DNA
36	qnorB gene, copy number, per unit sediment dry mass	qnorB/sed dm	#/g	Pastor, Ada	qnorB gene (encoding quinol-dependent nitric oxide reductase ) in sediment samples, quantified from extracted DNA

License:

Creative Commons Attribution-ShareAlike 4.0 International (CC-BY-SA-4.0)

Status:

Curation Level: Enhanced curation (CurationLevelC)

Size:

1477 data points

Data

Download dataset as tab-delimited text — use the following character encoding:

1 Event	2 Sample label	3 Latitude	4 Longitude	5 Date/Time	6 Elevation [m]	7 pH (streamwater)	8 Temp [°C] (streamwater)	9 EC [µS/cm] (streamwater)	10 [NH4]+ [mg/l] (streamwater)	11 [NO3]- [mg/l] (streamwater)	12 DOC [mg/l] (streamwater)	13 DON [mg/l] (streamwater)	14 TDN [mg/l] (streamwater)	15 pH (sediment)	16 EC [µS/cm] (sediment)	17 >2 mm [%] (Sediment size >2000 m)	18 2000-630 µm [%] (Sediment size 2000-630 m)	19 630-200 µm [%] (Sediment size 630-200 m)	20 200-63 µm fS [%] (Sediment size 200 63 m)	21 LOI [%]	22 [NH4]+ [mg/l] (sediment)	23 [NO3]- [mg/l] (sediment)	24 N [%] (sediment)	25 C [%] (sediment)	26 NDVI	27 Catch area [km²]	28 Slope [deg]	29 16S rRNA gene/sed dm [#/g]	30 AOA-amoA/sed dm [#/g] (Archaeal amoA gene (encoding ...)	31 AOB-amoA/sed dm [#/g] (Bacterial amoA gene (encoding...)	32 nifH/sed dm [#/g] (nifH gene (encoding the nitro...)	33 nrxB/sed dm [#/g] (nrxB gene (encoding the beta ...)	34 nosZ/sed dm [#/g] (nosZ gene (encoding nitrous o...)	35 nirS/sed dm [#/g] (nirS gene (encoding cytochrom...)	36 qnorB/sed dm [#/g] (qnorB gene (encoding quinol-d...)
Toolik_Inlet_microbes	Toolik Inlet	68.6251	-149.5927	2021-08-21	723	7.84	7.4	120.1	0.005	0.0340	4.00	0.18	0.21000	7.00	52.20	77.26	11.80	9.28	1.43	4.86	1.6400	0.0300	0.34	15.46	0.703	48.324808280	3.995	553335536.700	99736.214	176476.244	15850191.590	72532274.230	1290938.685	6933529.165	149494295.600
Imnavait_microbes	Imnavait	68.6250	-149.3247	2021-08-24	864	6.87	6.1	129.5	0.005	0.0438	10.32	0.29	0.34000	5.91	53.00	13.01	18.19	47.87	20.24	12.92	6.4700	0.7200	0.70	51.03	0.714	2.633832777	4.598	3091104417.000	52256.591	356835.518	365023814.000	173861530.300	11005134.510	12053298.290	525415902.700
Kuparuk_microbes	Kuparuk	68.6408	-149.3998	2021-08-25	748	6.89	6.1	90.4	0.010	0.0665	3.97	0.16	0.23000	6.79	26.90	63.56	14.80	15.01	6.03	4.47	3.2300	0.1400	0.36	18.66	0.715	133.544655900	7.121	200590613.800	51720.683	182177.717	5086780.717	41081632.430	816595.396	3928687.166	101031331.000
Oksrukuyik_microbes	Oksrukuyik	68.6858	-149.1003	2021-08-26	762	6.84	6.4	54.3	0.021	0.0399	5.98	0.21	0.27000	6.65	33.20	48.93	18.25	20.19	11.69	9.92	3.9900	0.2300	0.57	24.77	0.694	72.169192810	3.296	760132558.700	372374.748	117956.023	70691422.910	90490254.420	2906626.306	11902179.880	278148865.100
Trevor_microbes	Trevor	68.2834	-149.3716	2021-07-09	890	8.28	3.8	387.9	0.001	0.3360	0.54	0.16	0.49000	8.96	58.87	44.74	33.14	17.86	4.33	3.29	0.4400	0.2100	0.19	11.10	0.298	1.366519000	31.770	19395800.720	3606.666	3652.818	142123.088	1861226.982	69078.331	220289.266	9074835.912
Rengleng_River_microbes	Rengleng River	67.7533	-133.8625	2021-08-24	34	7.41	11.7	258.6	0.036	0.0284	7.59	0.29	0.35000	7.40	125.00	2.67	15.57	75.18	6.10	3.83	17.7600	0.0300	0.24	7.67	0.723	1065.099471000	1.513	233281325.800	24077.741	132362.681	19623144.280	9351845.259	498068.533	5866757.433	86712575.380
Caribou_Creek_microbes	Caribou Creek	68.0872	-133.4922	2021-08-24	20	7.16	10.6	244.0	0.016	0.0152	5.24	0.20	0.23000	7.43	42.70	7.74	25.63	56.33	9.67	2.62	6.2000	0.2200	0.19	6.58	0.690	574.612433200	1.469	12919505.760	2137.805	46147.861	7380829.356	1673143.259	239263.956	1427102.893	24697247.570
Havipak_Creek_microbes	Havipak Creek	68.3130	-133.5235	2021-08-19	48	7.13	5.3	160.6	0.005	0.0727	20.60	0.70	0.77000	8.16	147.60	81.14	11.64	6.33	0.87	6.03	12.8800	0.1000	0.42	37.36	0.757	20.530747460	3.061	695319232.400	36515.281	272544.724	35757249.860	38752798.570	1598344.199	10551889.830	181941338.000
Austre-Bayelva_microbes	Austre-Bayelva	78.9199	11.8262	2021-08-23	53	7.96	0.2	66.2	0.048	0.0204	0.70	0.04	0.11000	7.93	76.27	41.13	19.00	34.40	5.13	6.02	0.0500	0.5500	0.43	25.13	-0.068	13.022567330	17.080	590782565.600	4606116.307	64653.619	1013350.056	24670120.100	613146.902	264702.107	69369370.460
Mørebekken_microbes	Mørebekken	78.9363	11.8241	2021-08-25	49	7.98	3.7	159.1	0.002	0.0830	0.37	0.05	0.13000	8.71	56.63	62.51	18.56	16.18	2.25	1.68	0.4200	0.1100	0.16	28.87	-0.144	2.130487840	19.040	88108019.160	83698.134	30466.677	618546.883	8898070.845	242307.056	478764.841	26533286.600
Storvatenet_microbes	Storvatenet	78.9228	11.8860	2021-08-21	56	8.72	9.9	268.7	0.041	0.0206	2.81	0.20	0.26000	7.33	152.93	60.90	16.06	15.99	6.21	6.84	9.6300	0.2400	0.49	31.13	0.234	0.383243180	3.820	556576529.200	39408.148	494159.642	34820513.950	38940812.480	1295621.907	14689566.710	149247730.400
Bayelva_microbes	Bayelva	78.9203	11.7959	2021-08-27	50	8.25	3.7	123.7	0.001	0.0459	0.54	0.10	0.15000	8.16	153.10	0.98	21.32	63.09	13.86	2.05	2.0500	0.0900	0.14	66.12	0.032	0.224691040	8.790	705789965.500	17543.713	356356.959	40395814.810	12162276.530	1340552.606	3074538.474	137656594.400
Voitelva_microbes	Voitelva	78.9066	12.0075	2021-08-28	63	7.43	1.2	145.5	0.021	0.0382	0.48	0.05	0.11000	8.41	299.07	54.93	25.12	14.88	4.15	0.85	0.0600	0.0800	0.11	2.90	-0.173	4.031452590	23.720	994774.234	846.039	1283.465	38556.165	19623.798	8749.626		61082.218
Kærelv_A_microbes	Kærelv A July	74.4947	-20.4963	2021-07-10	190	7.70	10.6	101.0	0.006	0.0180	1.38	0.04	0.06600	7.87	97.00	46.85	28.41	21.75	2.06	2.03	0.1656	0.4328	0.15	2.81	0.160	0.855400251	13.550	18344569.290	402912.992	568419.311	781539.640	6459304.160	275443.320	424547.073	9170451.718
Kærelv_C_microbes	Kærelv C July	74.4710	-20.5188	2021-07-09	14	7.60	10.6	144.0	0.028	0.0030	1.71	0.07	0.10100	7.31	39.10	43.95	35.46	17.04	2.91	0.90	0.6970	0.3632	0.14	0.01	0.340	4.721680025	8.030	28426733.960	106578.226	43451.608	4396612.800	3048267.234	267685.073	704162.860	15048115.520
West_1_microbes	West 1 July	74.4739	-20.6076	2021-07-11	35	6.30	15.4	24.0	0.021	0.0060	1.37	0.07	0.09700	6.53	54.80	60.99	20.33	15.30	1.73	1.38	2.5540	0.4816	0.15	4.81	0.080	5.063849926	12.120	757354471.900	62861.242	156566.225	23987297.190	125522133.500	2379726.364	8061304.677	196302894.400
West_2_microbes	West 2 July	74.4716	-20.6556	2021-07-13	119	5.70	4.6	12.0	0.016	0.0160	0.58	0.03	0.06600	6.71	7.97	26.78	29.22	32.36	10.87	0.72	0.8072	0.3530	0.12	1.02	0.020	1.658958064	20.280	19425038.620	19577.487	112797.969	269204.093	5949691.733	363822.107		16125762.440
West_3_microbes	West 3 July	74.4605	-20.6927	2021-07-13	13	7.90	7.0	16.0	0.014	0.0240	0.34	0.01	0.05200	6.72	25.80	44.66	30.88	19.22	4.59	0.85	0.9255	0.2062	0.18	2.16	-0.010	4.772101450	21.550	90349483.890	103652.283	82344.506	680977.728	9938077.119	838387.075	111565.917	23753742.650
Tørvedammen_microbes	Tørvedammen July	74.4793	-20.5572	2021-07-14	37	6.50	12.0	59.0	0.016	0.0000	2.59	0.12	0.14100	6.87	63.90	58.97	31.72	8.13	0.88	1.43	4.1561	0.4171	0.16	2.59	0.430	1.371913631	4.410	113194470.800	4881.666	388532.365	6492322.236	9973914.845	800824.148	755191.894	28701540.620
Lindemann_tributary_microbes	Lindemann tributary July	74.5392	-20.6534	2021-07-15	182	7.80	4.5	985.0	0.028	0.0330	0.26	0.01	0.06700	7.74	280.00	18.48	18.92	54.85	6.97	0.92	0.2120	0.1312	0.13	1.87	0.020	0.483537044	24.010	4930627.654	14257.788	23488.206	507678.266	473260.022	72354.906	72299.848	1601814.704
Palnatoke_microbes	Palnatoke July	74.5273	-20.5752	2021-07-15	213	6.80	8.9	53.0	0.037	0.0250	0.48	0.01	0.07300	7.87	87.80	8.24	14.87	68.46	7.90	2.44	0.0396	0.1669	0.11	4.11	0.020	11.257110420	16.550	14514939.810	41118.934	172565.220	676976.841	1397861.667	295695.685	613907.190	5143333.805
Jurassic_1_microbes	Jurassic 1 July	74.4322	-20.2899	2021-07-17	59	4.30	8.4	705.0	0.093	0.0330	0.32	0.00	0.08900	4.89	122.50	15.13	2.74	63.68	17.84	2.71	0.3969	0.0977	0.13	9.82	0.030	10.244026840	16.710	4770991.693	10406.971	20365.744	2550783.162	172514.775	69218.168	75118.604	1362111.859
Jurassic_2_microbes	Jurassic 2 July	74.4398	-20.3080	2021-07-17	106	4.10	10.2	1316.0	0.091	0.1300	0.31	0.00	0.16900	4.31	282.00	40.26	28.57	28.77	2.04	6.77	0.3172	0.2479	0.22	22.92	0.080	1.336045326	7.110	6638960.788	35437.678	19740.483	36244035.990	103691.935	99062.265		1299834.375
Aucella_microbes	Aucella July	74.5033	-20.5418	2021-07-18	154	7.30	7.6	153.0	0.042	0.0590	0.21	0.00	0.08500	7.51	152.50	6.03	29.78	38.30	24.60	3.78	0.1887	0.1358	0.24	17.57	0.020	3.837075527	15.570	8392579.222	34336.021	237360.732	629186.308	748719.864	174976.547	326147.933	4459131.475
Grænseelv_microbes	Grænseelv July	74.4732	-20.5083	2021-07-19	23	6.90	13.2	212.0	0.035	0.0130	1.30	0.07	0.11500	7.16	59.40	26.93	25.90	38.68	8.28	0.81	0.2905	0.1446	0.12	1.24	0.370	2.447717749	8.170	70273631.660	186887.134	87828.878	2163691.170	6312544.103	284119.287	789199.193	13559154.740
Grænseelv_microbes	Grænseelv August	74.4732	-20.5083	2021-08-05	23	6.90	10.2	268.0	0.005	0.0050	2.22	0.21	0.22200	7.01	71.60	15.13	52.05	28.09	4.32	0.58	0.7514	0.1834	0.08	0.66	0.330	2.447717749	8.170	7484880.542	48822.386	38504.330	683722.138	3007506.596	132195.912	271036.768	8808239.564
Kærelv_C_microbes	Kærelv C August	74.4710	-20.5188	2021-08-05	14	7.20	11.5	304.0	0.013	0.0060	0.94	0.04	0.06300	6.91	45.80	23.33	55.22	19.92	1.30	0.85	0.2997	0.1807	0.00	0.04	0.310	4.721680025	8.030	111589163.100	106621.140	64603.011	2217326.163	7700482.215	259916.086	664820.563	15878052.800
Kærelv_A_microbes	Kærelv A August	74.4947	-20.4963	2021-08-07	190	7.50	8.7	313.0	0.068	0.0320	1.49	0.17	0.27300	8.00	125.30	5.52	34.44	57.31	2.01	2.84	0.9102	0.6418	0.10	0.75	0.140	0.855400251	13.550	218037759.600	555186.059	1754125.257	1248297.339	9949396.706	738794.767	1205877.447	21205710.650
Aucella_microbes	Aucella August	74.5033	-20.5418	2021-08-07	154	7.50	7.1	306.0	0.006	0.1310	0.60	0.10	0.24100	7.62	177.50	6.85	20.17	59.58	12.80	3.80	0.4667	0.2584	0.00	0.01	0.040	3.837075527	15.570	4248454.995	1944.316	53483.958	257975.039	77735.367	56340.192	111012.337	801136.688
Palnatoke_microbes	Palnatoke August	74.5273	-20.5752	2021-08-09	213	7.00	9.5	86.0	0.017	0.0320	0.38	0.03	0.08000	7.64	136.40	20.10	8.46	45.21	25.84	2.88	0.8254	0.1270	0.02	0.24	0.030	11.257110420	16.550	4096565.744	3366.211	43936.922	395929.521	118987.138	82519.052	133819.202	621620.652
Lindemann_tributary_microbes	Lindemann tributary August	74.5392	-20.6534	2021-08-09	182	7.80	11.9	1846.0	0.025	0.0080	0.63	0.05	0.08800	7.88	231.00	6.73	16.63	59.47	16.48	1.49	0.1326	0.1440	0.01	0.12	0.040	0.483537044	24.010	8258002.144	10753.227	31359.097	277992.023	301789.120	92382.036	80799.448	2390870.317
Tørvedammen_microbes	Tørvedammen August	74.4793	-20.5572	2021-08-11	37	6.00	9.4	107.0	0.013	0.0320	2.96	0.24	0.28300	6.70	128.00	61.38	26.98	10.42	0.89	0.98	0.3189	0.2071	0.00	0.07	0.410	1.371913631	4.410	112406772.900	8796.974	46311.765	13994961.400	3911994.534	386889.641	445795.712	24574757.560
West_1_microbes	West 1 August	74.4739	-20.6076	2021-08-11	35	7.20	13.4	54.0	0.030	0.0140	1.64	0.08	0.12400	6.51	139.70	41.59	44.86	11.77	1.29	0.78	0.2740	0.3388	0.00	0.04	0.070	5.063849926	12.120	147945480.600	4508.816	51500.383	3159323.841	19596110.110	509692.885	2140275.523	36163659.000
West_4_microbes	West 4 August	74.4486	-20.7854	2021-08-12	9	7.10	9.3	161.0	0.020	0.1490	0.89	0.06	0.22900	0.17	20.60	32.25	24.94	32.16	9.55	0.57	0.5088	0.1310	0.00	0.03	0.070	0.434486684	28.200	27299731.860	244440.199	13478.425	315446.892	4947014.443	184264.280	58460.846	20303500.770
West_5_microbes	West 5 August	74.4542	-20.8525	2021-08-12	6	7.20	6.1	80.0	0.025	0.0610	1.44	0.09	0.17700	7.14	21.60	66.24	21.40	10.28	2.13	0.98	1.3887	0.1829	0.07	0.95	0.050	0.750610638	28.650	249247780.900	61320.414	27502.691	1351421.509	35506286.380	872451.918	167452.490	79770308.610
Jurassic_2_microbes	Jurassic 2 August	74.4398	-20.3080	2021-08-14	106	3.70	9.3	4140.0	0.186	0.0910	0.76	0.00	0.22100	4.17	932.00	15.80	43.72	36.20	3.65	8.21	0.1754	0.2646	0.26	3.41	0.080	1.336045326	7.110	14465238.750	21029.566	5575.658	38391380.130	182952.901	95370.054		1254682.422
Jurassic_1_microbes	Jurassic 1 August	74.4322	-20.2899	2021-08-14	59	4.30	9.1	1347.0	0.092	0.0590	0.40	0.02	0.16800	3.69	237.00	2.72	6.49	48.16	41.35	3.60	0.1486	0.1274	0.00	0.02	0.040	10.244026840	16.710	7353398.335	6347.338	5192.299	2656521.585	42777.829	41562.835		479605.044
Kærelv_C_microbes	Kærelv C September	74.4710	-20.5188	2021-09-08	14	7.30	3.1	393.0	0.049	0.0080	1.80	0.05	0.10920	6.18	56.80	4.87	76.99	15.70	2.01	1.04	0.1948	0.3056	1.02	13.52	0.230	4.721680025	8.030	61986956.610	121164.350	66512.526	4380124.908	5206834.646	292506.236	1212844.436	17378797.090
Kærelv_A_microbes	Kærelv A September	74.4947	-20.4963	2021-09-08	190	7.50	3.2	530.0	0.029	0.2070	1.16	0.03	0.26970	7.50	142.90	34.28	38.29	24.71	2.46	2.92	0.0378	0.7508	0.15	1.98	0.110	0.855400251	13.550	236055686.100	706266.306	1033734.044	7663320.608	10618790.880	942062.626	1690229.041	21838159.270
West_5_microbes	West 5 September	74.4542	-20.8525	2021-09-10	6	7.10	1.3	133.0	0.034	0.3410	1.04	0.02	0.39660	6.84	146.10	53.29	9.10	16.95	20.20	8.08	58.2880	1.0188	0.49	6.50	0.050	0.750610638	28.650	4228805489.000	207975.272	798485.822	30967073.900	120539915.400	8162535.360	2380167.834	388917463.100
West_4_microbes	West 4 September	74.4486	-20.7854	2021-09-10	9	7.00	1.1	234.0	0.023	0.1600	1.14	0.04	0.22550	7.01	53.90	78.53	12.62	6.37	2.27	1.12	3.4762	0.6876	2.16	28.80	0.070	0.434486684	28.200	93911191.380	17756.829	18844.747	1400494.636	19258581.880	510235.129	139385.067	45076402.340
Aucella_microbes	Aucella September	74.5033	-20.5418	2021-09-12	154	7.70	0.0	1201.0	0.033	0.6090	0.68	0.06	0.69880	7.99	235.00	64.59	11.00	18.74	5.02	2.86	0.1511	0.1660	0.00	0.04	0.040	3.837075527	15.570	7538601.781	20227.284	39464.591	979456.679	388055.801	146265.247	435032.715	1932549.915
Tørvedammen_microbes	Tørvedammen September	74.4793	-20.5572	2021-09-12	37	7.60	0.0	81.0	0.020	0.0100	1.75	0.09	0.12260	6.94	31.10					2.93	0.3219	0.0947	0.00	0.03	0.290	1.371913631	4.410	1406481860.000	56675.614	177432.070	128555279.000	34454560.670	2721043.031	3745742.508	79552193.610
West_1_microbes	West 1 September	74.4739	-20.6076	2021-09-12	35	6.40	0.1	92.0	0.007	0.0210	2.30	0.10	0.12720	7.05	38.70	31.12	26.53	32.14	10.04	2.28	0.0382	0.1461	0.00	0.02	0.040	5.063849926	12.120	637871054.200	85006.306	83476.477	6825362.927	87156746.440	1739289.337	1875899.571	122281550.000
West_3_microbes	West 3 September	74.4605	-20.6927	2021-09-13	13	7.40	1.3	43.0	0.019	0.2200	0.22	0.00	0.22280	7.44	20.60	55.33	11.69	20.33	12.00	0.97	1.4377	0.4148	0.00	0.02	0.020	4.772101450	21.550	315648322.600	78887.456	116143.541	791395.656	10733243.030	1147757.588	200839.731	42265489.590
West_2_microbes	West 2 September	74.4716	-20.6556	2021-09-13	119	7.10	2.5	44.0	0.000	0.1310	0.67	0.01	0.13850	7.41	12.54	70.93	16.53	10.78	1.17	0.33	0.5675	0.4065	0.00	0.01	0.040	1.658958064	20.280	14957500.010	1323.446	21687.178	71436.546	1945662.426	188529.799		9745160.372
Grænseelv_microbes	Grænseelv September	74.4732	-20.5083	2021-09-15	23	7.00	0.1	358.0	0.050	0.0890	2.07	0.04	0.18140	7.48	105.00	21.29	43.40	30.68	4.05	0.67	0.0535	0.3452	0.00	0.05	0.260	2.447717749	8.170	144870530.600	191206.861	77506.365	2751635.997	4951581.328	500879.276	1584577.365	23404822.970
Zackenberg_River_microbes	Zackenberg River September	74.4699	-20.5742	2021-09-17	30	7.40	0.0	1131.0	0.003	0.0540	6.78	0.04	0.09536	7.57	50.50	66.73	20.36	11.13	1.80	0.31	0.2153	0.2065	0.00	0.02		482.220707800	16.040	3317165.337	15485.574	8646.283	60566.245	616818.965	38802.523	99444.878	1721330.832