Citation:

PANGAEA.918920

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Abstract:

Here we present data from food selectivity experiments of four abundant Alpine decomposer species (two earthworms and two millipedes). We conducted cafeteria feeding experiments in microcosms in a climate chamber offering to the specimens three representative alpine litter types of differing in litter quality (i.e. dwarf shrub, grass, and forb) as single litters and mixtures. We monitored parameters such as biomass, consumption rates (absolute and relative), and calculated from the latter food selectivity indices to investigate preferences. The tested specimens showed a high intra- and interspecific variability as they fed on all offered litter types and qualities. The epi-endogeic earthworm Lumbricus rubellus (Hoffmeister, 1843), dominant in alpine pastureland, had highest mean consumption rates (34.7 ± 24.7 mg), while Dendrobaena octaedra (Savigny, 1826), a strictly epigeic earthworm, showed considerably lower consumption rates. The tested millipedes showed similar consumption patterns, with Cylindroiulus fulviceps (Latzel, 1884) being less active than C. meinerti (Verhoeff, 1891). We found several cases where litter mixtures were preferred over single litters, including higher consumption rates of low-quality dwarf shrub litter by D. octaedra and C. meinerti when mixed with high-quality litter.

Nowadays, alpine pastureland are threatened mainly by abandonment which reverted to shrubland. Our results helped us to understand that alpine soil macro-decomposer such as earthworms and millipedes are generalists feeding on a wide range of litter qualities, allowing them, in theory, to better adapt to the new evolving litter resources that become available when pastures are abandoned.

Related to:

Steinwandter, Michael; Seeber, Julia (2020): The buffet is open: Alpine soil macro-decomposers feed on a wide range of litter types in a microcosm cafeteria experiment. Soil Biology and Biochemistry, 144, 107786, https://doi.org/10.1016/j.soilbio.2020.107786

Parameter(s):

#	Name	Short Name	Unit	Principal Investigator	Comment
1	Sample ID	Sample ID		Steinwandter, Michael
2	Species	Species		Steinwandter, Michael
3	Treatment: food	T:food		Steinwandter, Michael
4	Biomass	Biom	mg	Steinwandter, Michael	Biomass at day 0
5	Biomass	Biom	mg	Steinwandter, Michael	Biomass at day 5
6	Biomass	Biom	mg	Steinwandter, Michael	Biomass at day 10
7	Litter consumption	Litter cons	mg	Steinwandter, Michael
8	Litter consumption	Litter cons	%	Steinwandter, Michael
9	Manly's alpha index	Manlys alpha		Steinwandter, Michael
10	Ivlev's electivity index	Ivlevs electrivity		Steinwandter, Michael
11	Forage ratio	Forage		Steinwandter, Michael

License:

Creative Commons Attribution-NonCommercial 4.0 International (CC-BY-NC-4.0)

Size:

3080 data points

Data

Download dataset as tab-delimited text — use the following character encoding:

1 Sample ID	2 Species	3 T:food	4 Biom [mg] (Biomass at day 0)	5 Biom [mg] (Biomass at day 5)	6 Biom [mg] (Biomass at day 10)	7 Litter cons [mg]	8 Litter cons [%]	9 Manlys alpha	10 Ivlevs electrivity	11 Forage
LR_01	Lumbricus rubellus	shrub	1294.13	1096.17	1156.650	18.304	5.160	3.366	-0.524	0.273
LR_02	Lumbricus rubellus	shrub	1875.60	1352.01	1357.840	20.176	5.915	3.167	-0.480	0.332
LR_03	Lumbricus rubellus	shrub	1367.55	1106.80	1131.984	92.728	27.398	16.278	0.101	1.180
LR_04	Lumbricus rubellus	shrub	1682.38	925.27	1078.830	29.120	8.826	20.975	0.169	1.458
LR_05	Lumbricus rubellus	shrub	1178.83	1039.52	1126.990	20.176	5.789	4.711	-0.449	0.372
LR_06	Lumbricus rubellus	shrub	1471.69	1254.12	1286.050	26.696	8.348	10.270	-0.131	0.740
LR_07	Lumbricus rubellus	shrub	1487.32	1186.70	1343.840	17.358	5.123	6.063	-0.339	0.460
LR_08	Lumbricus rubellus	shrub	1149.42	838.46	984.780	28.956	9.715	9.678	-0.119	0.726
LR_09	Lumbricus rubellus	shrub	1298.29	1269.12	1286.300	19.372	5.759	4.446	-0.468	0.348
LR_10	Lumbricus rubellus	shrub	1049.48	1095.82	907.560	52.812	16.182	9.658	-0.120	0.784
LR_01	Lumbricus rubellus	grass	1294.13	1096.17	1156.650	80.774	38.242	30.615	0.396	2.027
LR_02	Lumbricus rubellus	grass	1875.60	1352.01	1357.840	3.162	1.497	0.783	-0.837	0.084
LR_03	Lumbricus rubellus	grass	1367.55	1106.80	1131.984	6.040	3.517	1.820	-0.728	0.151
LR_04	Lumbricus rubellus	grass	1682.38	925.27	1078.830	13.322	6.658	15.639	0.030	1.100
LR_05	Lumbricus rubellus	grass	1178.83	1039.52	1126.990	27.892	14.153	12.055	-0.036	0.910
LR_06	Lumbricus rubellus	grass	1471.69	1254.12	1286.050	17.070	9.445	11.689	-0.070	0.837
LR_07	Lumbricus rubellus	grass	1487.32	1186.70	1343.840	22.032	11.243	13.752	0.040	1.009
LR_08	Lumbricus rubellus	grass	1149.42	838.46	984.780	19.702	10.223	10.212	-0.094	0.764
LR_09	Lumbricus rubellus	grass	1298.29	1269.12	1286.300	22.668	12.437	9.954	-0.122	0.752
LR_10	Lumbricus rubellus	grass	1049.48	1095.82	907.560	18.968	10.704	6.194	-0.316	0.519
LR_01	Lumbricus rubellus	forb	1294.13	1096.17	1156.650	70.342	35.823	28.175	0.368	1.898
LR_02	Lumbricus rubellus	forb	1875.60	1352.01	1357.840	157.620	75.393	72.839	0.635	4.229
LR_03	Lumbricus rubellus	forb	1367.55	1106.80	1131.984	49.612	27.445	16.310	0.102	1.182
LR_04	Lumbricus rubellus	forb	1682.38	925.27	1078.830	0.084	0.044	0.099	-0.986	0.007
LR_05	Lumbricus rubellus	forb	1178.83	1039.52	1126.990	100.824	41.630	42.528	0.464	2.676
LR_06	Lumbricus rubellus	forb	1471.69	1254.12	1286.050	39.270	17.543	22.726	0.235	1.555
LR_07	Lumbricus rubellus	forb	1487.32	1186.70	1343.840	65.636	31.984	44.444	0.510	2.871
LR_08	Lumbricus rubellus	forb	1149.42	838.46	984.780	62.920	34.858	40.586	0.477	2.604
LR_09	Lumbricus rubellus	forb	1298.29	1269.12	1286.300	28.142	12.154	9.713	-0.133	0.735
LR_10	Lumbricus rubellus	forb	1049.48	1095.82	907.560	72.838	33.544	22.358	0.239	1.626
LR_01	Lumbricus rubellus	shrub + grass	1294.13	1096.17	1156.650	18.062	7.905	5.231	-0.353	0.419
LR_02	Lumbricus rubellus	shrub + grass	1875.60	1352.01	1357.840	23.182	12.949	7.204	-0.131	0.726
LR_03	Lumbricus rubellus	shrub + grass	1367.55	1106.80	1131.984	82.396	33.510	20.748	0.199	1.443
LR_04	Lumbricus rubellus	shrub + grass	1682.38	925.27	1078.830	13.468	8.419	19.964	0.146	1.391
LR_05	Lumbricus rubellus	shrub + grass	1178.83	1039.52	1126.990	47.550	22.178	19.807	0.186	1.426
LR_06	Lumbricus rubellus	shrub + grass	1471.69	1254.12	1286.050	35.944	18.333	23.860	0.256	1.625
LR_07	Lumbricus rubellus	shrub + grass	1487.32	1186.70	1343.840	5.754	2.605	3.043	-0.599	0.234
LR_08	Lumbricus rubellus	shrub + grass	1149.42	838.46	984.780	36.332	16.990	17.633	0.159	1.269
LR_09	Lumbricus rubellus	shrub + grass	1298.29	1269.12	1286.300	6.856	3.359	2.561	-0.651	0.203
LR_10	Lumbricus rubellus	shrub + grass	1049.48	1095.82	907.560	125.600	57.059	46.252	0.470	2.766
LR_01	Lumbricus rubellus	shrub + forb	1294.13	1096.17	1156.650	43.148	29.936	22.600	0.288	1.587
LR_02	Lumbricus rubellus	shrub + forb	1875.60	1352.01	1357.840	7.752	4.278	2.271	-0.595	0.240
LR_03	Lumbricus rubellus	shrub + forb	1367.55	1106.80	1131.984	65.092	45.772	31.112	0.343	1.970
LR_04	Lumbricus rubellus	shrub + forb	1682.38	925.27	1078.830	7.368	4.594	10.675	-0.154	0.759
LR_05	Lumbricus rubellus	shrub + forb	1178.83	1039.52	1126.990	11.326	6.734	5.507	-0.387	0.433
LR_06	Lumbricus rubellus	shrub + forb	1471.69	1254.12	1286.050	13.486	8.089	9.937	-0.147	0.717
LR_07	Lumbricus rubellus	shrub + forb	1487.32	1186.70	1343.840	18.182	10.828	13.215	0.021	0.972
LR_08	Lumbricus rubellus	shrub + forb	1149.42	838.46	984.780	22.898	13.687	13.938	0.052	1.023
LR_09	Lumbricus rubellus	shrub + forb	1298.29	1269.12	1286.300	32.168	16.310	13.346	0.013	0.986
LR_10	Lumbricus rubellus	shrub + forb	1049.48	1095.82	907.560	24.954	13.447	7.901	-0.210	0.652
LR_01	Lumbricus rubellus	grass + forb	1294.13	1096.17	1156.650	8.618	3.924	2.543	-0.616	0.208
LR_02	Lumbricus rubellus	grass + forb	1875.60	1352.01	1357.840	27.024	12.435	6.898	-0.151	0.697
LR_03	Lumbricus rubellus	grass + forb	1367.55	1106.80	1131.984	42.128	17.668	9.884	-0.118	0.761
LR_04	Lumbricus rubellus	grass + forb	1682.38	925.27	1078.830	20.752	9.053	21.541	0.182	1.496
LR_05	Lumbricus rubellus	grass + forb	1178.83	1039.52	1126.990	13.580	5.491	4.461	-0.470	0.353
LR_06	Lumbricus rubellus	grass + forb	1471.69	1254.12	1286.050	28.024	13.141	16.599	0.095	1.165
LR_07	Lumbricus rubellus	grass + forb	1487.32	1186.70	1343.840	20.910	8.267	9.950	-0.113	0.742
LR_08	Lumbricus rubellus	grass + forb	1149.42	838.46	984.780	7.742	3.953	3.819	-0.515	0.295
LR_09	Lumbricus rubellus	grass + forb	1298.29	1269.12	1286.300	64.910	36.108	33.578	0.389	2.182
LR_10	Lumbricus rubellus	grass + forb	1049.48	1095.82	907.560	12.542	6.031	3.403	-0.547	0.292
LR_01	Lumbricus rubellus	shrub + grass + forb	1294.13	1096.17	1156.650	31.240	11.094	7.470	-0.197	0.588
LR_02	Lumbricus rubellus	shrub + grass + forb	1875.60	1352.01	1357.840	36.348	12.331	6.837	-0.155	0.692
LR_03	Lumbricus rubellus	shrub + grass + forb	1367.55	1106.80	1131.984	20.796	7.291	3.849	-0.509	0.314
LR_04	Lumbricus rubellus	shrub + grass + forb	1682.38	925.27	1078.830	14.048	4.775	11.106	-0.135	0.789
LR_05	Lumbricus rubellus	shrub + grass + forb	1178.83	1039.52	1126.990	31.070	12.924	10.932	-0.082	0.831
LR_06	Lumbricus rubellus	shrub + grass + forb	1471.69	1254.12	1286.050	11.580	4.090	4.920	-0.453	0.362
LR_07	Lumbricus rubellus	shrub + grass + forb	1487.32	1186.70	1343.840	21.000	7.934	9.532	-0.133	0.712
LR_08	Lumbricus rubellus	shrub + grass + forb	1149.42	838.46	984.780	12.998	4.272	4.134	-0.486	0.319
LR_09	Lumbricus rubellus	shrub + grass + forb	1298.29	1269.12	1286.300	80.250	29.689	26.402	0.303	1.794
LR_10	Lumbricus rubellus	shrub + grass + forb	1049.48	1095.82	907.560	18.702	7.444	4.233	-0.469	0.361
DO_01	Dendrobaena octaedra	shrub	176.93	134.14	161.620	13.126	5.212	14.685	0.048	1.045
DO_02	Dendrobaena octaedra	shrub	180.34	173.88	154.360	2.340	0.982	3.767	-0.582	0.270
DO_03	Dendrobaena octaedra	shrub	171.72	157.72	165.650	16.952	7.365	18.198	0.152	1.307
DO_04	Dendrobaena octaedra	shrub	156.06	125.73	112.480	3.448	1.442	6.789	-0.332	0.495
DO_05	Dendrobaena octaedra	shrub	110.05	109.12	122.440	8.916	4.037	5.799	-0.349	0.453
DO_06	Dendrobaena octaedra	shrub	220.08	191.70	216.480	21.290	8.464	13.328	-0.014	0.961
DO_07	Dendrobaena octaedra	shrub	215.13	211.36	226.700	3.718	1.440	1.464	-0.772	0.122
DO_08	Dendrobaena octaedra	shrub	110.14	70.33	99.340	23.322	9.222	28.095	0.319	1.936
DO_09	Dendrobaena octaedra	shrub	124.06	124.58	128.420	3.816	1.128	3.306	-0.601	0.242
DO_10	Dendrobaena octaedra	shrub	98.51	113.46	109.620	26.694	10.992	31.673	0.365	2.170
DO_01	Dendrobaena octaedra	grass	176.93	134.14	161.620	9.050	4.598	12.914	-0.015	0.922
DO_02	Dendrobaena octaedra	grass	180.34	173.88	154.360	4.598	2.306	8.908	-0.234	0.634
DO_03	Dendrobaena octaedra	grass	171.72	157.72	165.650	3.004	1.434	3.436	-0.582	0.254
DO_04	Dendrobaena octaedra	grass	156.06	125.73	112.480	0.084	0.049	0.228	-0.967	0.017
DO_05	Dendrobaena octaedra	grass	110.05	109.12	122.440	4.614	2.590	3.692	-0.527	0.290
DO_06	Dendrobaena octaedra	grass	220.08	191.70	216.480	7.610	4.773	7.371	-0.292	0.542
DO_07	Dendrobaena octaedra	grass	215.13	211.36	226.700	6.340	3.654	3.758	-0.508	0.310
DO_08	Dendrobaena octaedra	grass	110.14	70.33	99.340	7.502	4.482	13.315	-0.031	0.941
DO_09	Dendrobaena octaedra	grass	124.06	124.58	128.420	0.000	0.000	0.000	-1.000	0.000
DO_10	Dendrobaena octaedra	grass	98.51	113.46	109.620	5.520	2.736	7.546	-0.302	0.540
DO_01	Dendrobaena octaedra	forb	176.93	134.14	161.620	8.708	2.444	6.788	-0.319	0.490
DO_02	Dendrobaena octaedra	forb	180.34	173.88	154.360	4.898	1.546	5.951	-0.412	0.425
DO_03	Dendrobaena octaedra	forb	171.72	157.72	165.650	7.260	2.274	5.472	-0.409	0.404
DO_04	Dendrobaena octaedra	forb	156.06	125.73	112.480	8.804	2.771	13.136	-0.019	0.951
DO_05	Dendrobaena octaedra	forb	110.05	109.12	122.440	7.704	2.302	3.277	-0.568	0.258
DO_06	Dendrobaena octaedra	forb	220.08	191.70	216.480	13.030	4.627	7.140	-0.306	0.525
DO_07	Dendrobaena octaedra	forb	215.13	211.36	226.700	7.656	2.819	2.887	-0.598	0.239
DO_08	Dendrobaena octaedra	forb	110.14	70.33	99.340	5.842	2.169	6.367	-0.374	0.455
DO_09	Dendrobaena octaedra	forb	124.06	124.58	128.420	1.054	0.403	1.176	-0.837	0.087
DO_10	Dendrobaena octaedra	forb	98.51	113.46	109.620	1.268	0.485	1.323	-0.826	0.096
DO_01	Dendrobaena octaedra	shrub + grass	176.93	134.14	161.620	7.798	3.243	9.044	-0.187	0.650
DO_02	Dendrobaena octaedra	shrub + grass	180.34	173.88	154.360	9.176	4.171	16.270	0.058	1.147
DO_03	Dendrobaena octaedra	shrub + grass	171.72	157.72	165.650	11.010	4.715	11.489	-0.070	0.837
DO_04	Dendrobaena octaedra	shrub + grass	156.06	125.73	112.480	26.124	12.791	63.990	0.633	4.389
DO_05	Dendrobaena octaedra	shrub + grass	110.05	109.12	122.440	22.216	9.581	14.172	0.068	1.075
DO_06	Dendrobaena octaedra	shrub + grass	220.08	191.70	216.480	51.058	22.700	38.803	0.446	2.577
DO_07	Dendrobaena octaedra	shrub + grass	215.13	211.36	226.700	8.378	2.856	2.925	-0.594	0.242
DO_08	Dendrobaena octaedra	shrub + grass	110.14	70.33	99.340	13.132	5.924	17.731	0.108	1.243
DO_09	Dendrobaena octaedra	shrub + grass	124.06	124.58	128.420	31.614	13.557	42.444	0.499	2.913
DO_10	Dendrobaena octaedra	shrub + grass	98.51	113.46	109.620	13.108	5.564	15.570	0.043	1.099
DO_01	Dendrobaena octaedra	shrub + forb	176.93	134.14	161.620	3.792	1.710	4.731	-0.469	0.343
DO_02	Dendrobaena octaedra	shrub + forb	180.34	173.88	154.360	7.424	2.787	10.793	-0.143	0.766
DO_03	Dendrobaena octaedra	shrub + forb	171.72	157.72	165.650	1.560	0.638	1.521	-0.790	0.113
DO_04	Dendrobaena octaedra	shrub + forb	156.06	125.73	112.480	0.754	0.360	1.685	-0.778	0.123
DO_05	Dendrobaena octaedra	shrub + forb	110.05	109.12	122.440	3.064	1.212	1.715	-0.747	0.136
DO_06	Dendrobaena octaedra	shrub + forb	220.08	191.70	216.480	3.860	1.519	2.306	-0.703	0.172
DO_07	Dendrobaena octaedra	shrub + forb	215.13	211.36	226.700	94.384	41.877	54.772	0.578	3.548
DO_08	Dendrobaena octaedra	shrub + forb	110.14	70.33	99.340	2.728	1.121	3.274	-0.619	0.235
DO_09	Dendrobaena octaedra	shrub + forb	124.06	124.58	128.420	2.444	0.999	2.925	-0.639	0.215
DO_10	Dendrobaena octaedra	shrub + forb	98.51	113.46	109.620	13.214	6.156	17.281	0.093	1.216
DO_01	Dendrobaena octaedra	grass + forb	176.93	134.14	161.620	5.830	3.400	9.488	-0.164	0.682
DO_02	Dendrobaena octaedra	grass + forb	180.34	173.88	154.360	9.914	4.363	17.035	0.080	1.200
DO_03	Dendrobaena octaedra	grass + forb	171.72	157.72	165.650	7.888	4.013	9.743	-0.150	0.712
DO_04	Dendrobaena octaedra	grass + forb	156.06	125.73	112.480	7.072	2.824	13.392	-0.009	0.969
DO_05	Dendrobaena octaedra	grass + forb	110.05	109.12	122.440	72.742	34.140	58.770	0.607	3.829
DO_06	Dendrobaena octaedra	grass + forb	220.08	191.70	216.480	16.986	8.942	14.118	0.013	1.015
DO_07	Dendrobaena octaedra	grass + forb	215.13	211.36	226.700	11.660	4.952	5.127	-0.387	0.420
DO_08	Dendrobaena octaedra	grass + forb	110.14	70.33	99.340	7.800	3.438	10.160	-0.162	0.722
DO_09	Dendrobaena octaedra	grass + forb	124.06	124.58	128.420	15.454	8.116	24.659	0.283	1.744
DO_10	Dendrobaena octaedra	grass + forb	98.51	113.46	109.620	6.202	3.097	8.557	-0.245	0.612
DO_01	Dendrobaena octaedra	shrub + grass + forb	176.93	134.14	161.620	29.596	14.304	42.349	0.503	2.868
DO_02	Dendrobaena octaedra	shrub + grass + forb	180.34	173.88	154.360	26.938	9.300	37.276	0.429	2.558
DO_03	Dendrobaena octaedra	shrub + grass + forb	171.72	157.72	165.650	42.158	19.006	50.141	0.556	3.373
DO_04	Dendrobaena octaedra	shrub + grass + forb	156.06	125.73	112.480	0.380	0.166	0.779	-0.891	0.057
DO_05	Dendrobaena octaedra	shrub + grass + forb	110.05	109.12	122.440	19.092	8.548	12.575	0.011	0.959
DO_06	Dendrobaena octaedra	shrub + grass + forb	220.08	191.70	216.480	25.992	10.628	16.934	0.099	1.207
DO_07	Dendrobaena octaedra	shrub + grass + forb	215.13	211.36	226.700	56.784	25.021	29.068	0.381	2.120
DO_08	Dendrobaena octaedra	shrub + grass + forb	110.14	70.33	99.340	17.214	6.996	21.059	0.190	1.468
DO_09	Dendrobaena octaedra	shrub + grass + forb	124.06	124.58	128.420	24.172	8.377	25.489	0.298	1.800
DO_10	Dendrobaena octaedra	shrub + grass + forb	98.51	113.46	109.620	16.012	6.420	18.049	0.114	1.268
CM_01	Cylindroiulus meinerti	shrub	200.68	207.71	206.380	15.372	8.068	12.943	-0.061	0.913
CM_02	Cylindroiulus meinerti	shrub	135.72	142.81	142.530	18.352	10.320	10.079	-0.158	0.735
CM_03	Cylindroiulus meinerti	shrub	172.69	176.05	177.620	47.478	31.354	21.182	0.175	1.461
CM_04	Cylindroiulus meinerti	shrub	164.50	171.56	172.640	10.498	6.379	5.805	-0.405	0.445
CM_05	Cylindroiulus meinerti	shrub	125.36	128.61	127.680	36.566	18.017	25.315	0.237	1.721
CM_06	Cylindroiulus meinerti	shrub	127.23	134.93	132.110	41.562	20.167	23.399	0.194	1.585
CM_07	Cylindroiulus meinerti	shrub	86.25	92.45	92.770	29.114	16.244	18.603	0.108	1.295
CM_08	Cylindroiulus meinerti	shrub	170.37	172.80	172.710	9.736	5.623	4.824	-0.463	0.365
CM_09	Cylindroiulus meinerti	shrub	177.01	182.17	184.040	20.436	11.282	8.521	-0.234	0.636
CM_10	Cylindroiulus meinerti	shrub	96.33	98.99	101.300	18.006	10.233	13.602	-0.025	0.963
CM_01	Cylindroiulus meinerti	grass	200.68	207.71	206.380	8.226	7.930	12.713	-0.069	0.897
CM_02	Cylindroiulus meinerti	grass	135.72	142.81	142.530	17.918	15.920	16.045	0.058	1.134
CM_03	Cylindroiulus meinerti	grass	172.69	176.05	177.620	8.412	7.673	4.495	-0.483	0.358
CM_04	Cylindroiulus meinerti	grass	164.50	171.56	172.640	9.508	9.684	8.971	-0.218	0.675
CM_05	Cylindroiulus meinerti	grass	125.36	128.61	127.680	6.814	7.168	9.477	-0.216	0.685
CM_06	Cylindroiulus meinerti	grass	127.23	134.93	132.110	10.608	9.538	10.414	-0.176	0.750
CM_07	Cylindroiulus meinerti	grass	86.25	92.45	92.770	8.006	7.851	8.581	-0.250	0.626
CM_08	Cylindroiulus meinerti	grass	170.37	172.80	172.710	9.910	8.780	7.660	-0.272	0.569
CM_09	Cylindroiulus meinerti	grass	177.01	182.17	184.040	21.168	17.935	14.069	-0.007	1.011
CM_10	Cylindroiulus meinerti	grass	96.33	98.99	101.300	8.872	8.375	11.021	-0.124	0.788
CM_01	Cylindroiulus meinerti	forb	200.68	207.71	206.380	24.542	12.560	20.651	0.159	1.421
CM_02	Cylindroiulus meinerti	forb	135.72	142.81	142.530	46.216	28.242	30.709	0.331	2.012
CM_03	Cylindroiulus meinerti	forb	172.69	176.05	177.620	12.582	8.084	4.746	-0.463	0.377
CM_04	Cylindroiulus meinerti	forb	164.50	171.56	172.640	15.588	10.602	9.871	-0.174	0.739
CM_05	Cylindroiulus meinerti	forb	125.36	128.61	127.680	30.280	18.049	25.365	0.238	1.724
CM_06	Cylindroiulus meinerti	forb	127.23	134.93	132.110	15.344	9.452	10.316	-0.181	0.743
CM_07	Cylindroiulus meinerti	forb	86.25	92.45	92.770	15.308	8.458	9.274	-0.214	0.674
CM_08	Cylindroiulus meinerti	forb	170.37	172.80	172.710	31.326	19.169	17.738	0.111	1.243
CM_09	Cylindroiulus meinerti	forb	177.01	182.17	184.040	41.970	20.670	16.481	0.064	1.165
CM_10	Cylindroiulus meinerti	forb	96.33	98.99	101.300	24.852	15.323	20.957	0.175	1.441
CM_01	Cylindroiulus meinerti	shrub + grass	200.68	207.71	206.380	8.616	6.383	10.148	-0.176	0.722
CM_02	Cylindroiulus meinerti	shrub + grass	135.72	142.81	142.530	18.754	11.656	11.468	-0.098	0.830
CM_03	Cylindroiulus meinerti	shrub + grass	172.69	176.05	177.620	23.088	16.380	10.072	-0.147	0.763
CM_04	Cylindroiulus meinerti	shrub + grass	164.50	171.56	172.640	21.358	15.851	15.200	0.025	1.105
CM_05	Cylindroiulus meinerti	shrub + grass	125.36	128.61	127.680	8.904	5.699	7.478	-0.322	0.544
CM_06	Cylindroiulus meinerti	shrub + grass	127.23	134.93	132.110	11.690	9.708	10.610	-0.168	0.763
CM_07	Cylindroiulus meinerti	shrub + grass	86.25	92.45	92.770	8.108	6.094	6.599	-0.364	0.486
CM_08	Cylindroiulus meinerti	shrub + grass	170.37	172.80	172.710	16.666	12.057	10.709	-0.120	0.782
CM_09	Cylindroiulus meinerti	shrub + grass	177.01	182.17	184.040	19.880	15.701	12.157	-0.073	0.885
CM_10	Cylindroiulus meinerti	shrub + grass	96.33	98.99	101.300	9.212	6.563	8.553	-0.242	0.617
CM_01	Cylindroiulus meinerti	shrub + forb	200.68	207.71	206.380	19.640	11.782	19.289	0.128	1.333
CM_02	Cylindroiulus meinerti	shrub + forb	135.72	142.81	142.530	26.248	16.031	16.168	0.061	1.142
CM_03	Cylindroiulus meinerti	shrub + forb	172.69	176.05	177.620	65.000	42.708	31.361	0.319	1.990
CM_04	Cylindroiulus meinerti	shrub + forb	164.50	171.56	172.640	61.598	36.881	40.526	0.420	2.570
CM_05	Cylindroiulus meinerti	shrub + forb	125.36	128.61	127.680	18.210	8.946	11.943	-0.108	0.855
CM_06	Cylindroiulus meinerti	shrub + forb	127.23	134.93	132.110	33.196	19.522	22.563	0.178	1.534
CM_07	Cylindroiulus meinerti	shrub + forb	86.25	92.45	92.770	27.648	17.571	20.279	0.147	1.401
CM_08	Cylindroiulus meinerti	shrub + forb	170.37	172.80	172.710	22.262	13.297	11.892	-0.071	0.862
CM_09	Cylindroiulus meinerti	shrub + forb	177.01	182.17	184.040	59.042	35.332	31.027	0.321	1.992
CM_10	Cylindroiulus meinerti	shrub + forb	96.33	98.99	101.300	16.248	9.162	12.108	-0.080	0.862
CM_01	Cylindroiulus meinerti	grass + forb	200.68	207.71	206.380	9.994	7.979	12.794	-0.066	0.903
CM_02	Cylindroiulus meinerti	grass + forb	135.72	142.81	142.530	11.558	8.267	7.985	-0.263	0.589
CM_03	Cylindroiulus meinerti	grass + forb	172.69	176.05	177.620	22.486	17.751	11.002	-0.108	0.827
CM_04	Cylindroiulus meinerti	grass + forb	164.50	171.56	172.640	15.030	12.396	11.656	-0.098	0.864
CM_05	Cylindroiulus meinerti	grass + forb	125.36	128.61	127.680	8.934	7.529	9.974	-0.192	0.719
CM_06	Cylindroiulus meinerti	grass + forb	127.23	134.93	132.110	8.946	8.686	9.440	-0.221	0.683
CM_07	Cylindroiulus meinerti	grass + forb	86.25	92.45	92.770	12.168	9.552	10.536	-0.156	0.761
CM_08	Cylindroiulus meinerti	grass + forb	170.37	172.80	172.710	40.324	29.088	28.651	0.310	1.886
CM_09	Cylindroiulus meinerti	grass + forb	177.01	182.17	184.040	9.900	7.533	5.575	-0.414	0.425
CM_10	Cylindroiulus meinerti	grass + forb	96.33	98.99	101.300	9.148	7.068	9.235	-0.207	0.665
CM_01	Cylindroiulus meinerti	shrub + grass + forb	200.68	207.71	206.380	10.810	7.179	11.463	-0.118	0.812
CM_02	Cylindroiulus meinerti	shrub + grass + forb	135.72	142.81	142.530	10.864	7.832	7.547	-0.288	0.558
CM_03	Cylindroiulus meinerti	shrub + grass + forb	172.69	176.05	177.620	33.020	26.250	17.143	0.087	1.223
CM_04	Cylindroiulus meinerti	shrub + grass + forb	164.50	171.56	172.640	10.792	8.653	7.971	-0.271	0.603
CM_05	Cylindroiulus meinerti	shrub + grass + forb	125.36	128.61	127.680	11.484	7.872	10.449	-0.171	0.752
CM_06	Cylindroiulus meinerti	shrub + grass + forb	127.23	134.93	132.110	17.684	11.982	13.259	-0.064	0.942
CM_07	Cylindroiulus meinerti	shrub + grass + forb	86.25	92.45	92.770	36.112	22.041	26.129	0.255	1.757
CM_08	Cylindroiulus meinerti	shrub + grass + forb	170.37	172.80	172.710	29.448	19.930	18.526	0.130	1.292
CM_09	Cylindroiulus meinerti	shrub + grass + forb	177.01	182.17	184.040	24.618	15.717	12.171	-0.073	0.886
CM_10	Cylindroiulus meinerti	shrub + grass + forb	96.33	98.99	101.300	24.306	17.686	24.524	0.244	1.664
CF_01	Cylindroiulus fulviceps	shrub	135.81	145.49	148.800	36.674	24.475	18.468	0.139	1.364
CF_02	Cylindroiulus fulviceps	shrub	124.42	126.75	125.820	3.802	2.483	7.694	-0.291	0.545
CF_03	Cylindroiulus fulviceps	shrub	167.88	173.27	175.290	5.098	3.521	10.924	-0.136	0.770
CF_04	Cylindroiulus fulviceps	shrub	112.61	115.11	117.940	3.280	2.086	6.009	-0.399	0.428
CF_05	Cylindroiulus fulviceps	shrub	115.87	118.84	121.840	4.410	2.389	3.048	-0.581	0.247
CF_06	Cylindroiulus fulviceps	shrub	127.66	131.93	141.340	16.224	9.561	25.240	0.254	1.736
CF_07	Cylindroiulus fulviceps	shrub	151.78	154.77	152.430	3.194	1.937	1.014	-0.799	0.122
CF_08	Cylindroiulus fulviceps	shrub	95.49	93.18	95.560	5.394	3.229	10.885	-0.153	0.767
CF_09	Cylindroiulus fulviceps	shrub	159.05	159.16	166.500	3.940	2.378	5.312	-0.455	0.383
CF_10	Cylindroiulus fulviceps	shrub	145.35	141.87	143.390	3.036	1.748	3.589	-0.534	0.268
CF_01	Cylindroiulus fulviceps	grass	135.81	145.49	148.800	5.332	5.146	3.476	-0.565	0.287
CF_02	Cylindroiulus fulviceps	grass	124.42	126.75	125.820	5.006	5.023	15.769	0.052	1.103
CF_03	Cylindroiulus fulviceps	grass	167.88	173.27	175.290	4.944	4.505	14.049	-0.014	0.985
CF_04	Cylindroiulus fulviceps	grass	112.61	115.11	117.940	5.594	4.989	14.590	0.013	1.024
CF_05	Cylindroiulus fulviceps	grass	115.87	118.84	121.840	5.208	6.007	7.810	-0.200	0.621
CF_06	Cylindroiulus fulviceps	grass	127.66	131.93	141.340	4.996	4.707	12.110	-0.094	0.855
CF_07	Cylindroiulus fulviceps	grass	151.78	154.77	152.430	5.534	6.041	3.230	-0.484	0.380
CF_08	Cylindroiulus fulviceps	grass	95.49	93.18	95.560	3.660	3.384	11.417	-0.131	0.804
CF_09	Cylindroiulus fulviceps	grass	159.05	159.16	166.500	3.966	4.070	9.169	-0.219	0.655
CF_10	Cylindroiulus fulviceps	grass	145.35	141.87	143.390	22.736	22.194	51.088	0.588	3.399
CF_01	Cylindroiulus fulviceps	forb	135.81	145.49	148.800	15.934	10.461	7.270	-0.278	0.583
CF_02	Cylindroiulus fulviceps	forb	124.42	126.75	125.820	9.968	6.361	20.113	0.169	1.397
CF_03	Cylindroiulus fulviceps	forb	167.88	173.27	175.290	8.392	5.135	16.067	0.052	1.123
CF_04	Cylindroiulus fulviceps	forb	112.61	115.11	117.940	10.690	6.271	18.463	0.127	1.287
CF_05	Cylindroiulus fulviceps	forb	115.87	118.84	121.840	13.056	8.509	11.212	-0.029	0.880
CF_06	Cylindroiulus fulviceps	forb	127.66	131.93	141.340	11.176	6.832	17.773	0.091	1.240
CF_07	Cylindroiulus fulviceps	forb	151.78	154.77	152.430	128.906	79.917	83.199	0.643	5.026
CF_08	Cylindroiulus fulviceps	forb	95.49	93.18	95.560	7.456	4.211	14.268	-0.022	1.001
CF_09	Cylindroiulus fulviceps	forb	159.05	159.16	166.500	20.706	10.751	25.098	0.257	1.731
CF_10	Cylindroiulus fulviceps	forb	145.35	141.87	143.390	7.678	4.954	10.345	-0.075	0.759
CF_01	Cylindroiulus fulviceps	shrub + grass	135.81	145.49	148.800	4.088	3.149	2.105	-0.709	0.175
CF_02	Cylindroiulus fulviceps	shrub + grass	124.42	126.75	125.820	5.012	3.593	11.196	-0.115	0.789
CF_03	Cylindroiulus fulviceps	shrub + grass	167.88	173.27	175.290	4.212	3.078	9.528	-0.201	0.673
CF_04	Cylindroiulus fulviceps	shrub + grass	112.61	115.11	117.940	4.140	3.860	11.222	-0.115	0.792
CF_05	Cylindroiulus fulviceps	shrub + grass	115.87	118.84	121.840	6.730	4.157	5.353	-0.369	0.430
CF_06	Cylindroiulus fulviceps	shrub + grass	127.66	131.93	141.340	4.026	3.098	7.904	-0.295	0.562
CF_07	Cylindroiulus fulviceps	shrub + grass	151.78	154.77	152.430	4.518	3.334	1.758	-0.678	0.210
CF_08	Cylindroiulus fulviceps	shrub + grass	95.49	93.18	95.560	3.734	3.041	10.240	-0.183	0.722
CF_09	Cylindroiulus fulviceps	shrub + grass	159.05	159.16	166.500	6.812	5.057	11.451	-0.114	0.814
CF_10	Cylindroiulus fulviceps	shrub + grass	145.35	141.87	143.390	4.248	3.279	6.788	-0.274	0.502
CF_01	Cylindroiulus fulviceps	shrub + forb	135.81	145.49	148.800	61.990	32.407	25.769	0.273	1.806
CF_02	Cylindroiulus fulviceps	shrub + forb	124.42	126.75	125.820	6.222	4.666	14.622	0.015	1.025
CF_03	Cylindroiulus fulviceps	shrub + forb	167.88	173.27	175.290	10.402	6.313	19.875	0.154	1.380
CF_04	Cylindroiulus fulviceps	shrub + forb	112.61	115.11	117.940	6.482	3.910	11.369	-0.108	0.802
CF_05	Cylindroiulus fulviceps	shrub + forb	115.87	118.84	121.840	5.614	3.100	3.971	-0.488	0.321
CF_06	Cylindroiulus fulviceps	shrub + forb	127.66	131.93	141.340	7.778	4.702	12.096	-0.095	0.854
CF_07	Cylindroiulus fulviceps	shrub + forb	151.78	154.77	152.430	8.148	5.518	2.942	-0.518	0.347
CF_08	Cylindroiulus fulviceps	shrub + forb	95.49	93.18	95.560	13.614	6.737	23.129	0.210	1.601
CF_09	Cylindroiulus fulviceps	shrub + forb	159.05	159.16	166.500	5.698	3.614	8.121	-0.275	0.582
CF_10	Cylindroiulus fulviceps	shrub + forb	145.35	141.87	143.390	6.984	3.960	8.226	-0.185	0.606
CF_01	Cylindroiulus fulviceps	grass + forb	135.81	145.49	148.800	49.690	45.398	39.811	0.421	2.529
CF_02	Cylindroiulus fulviceps	grass + forb	124.42	126.75	125.820	7.208	5.557	17.495	0.102	1.220
CF_03	Cylindroiulus fulviceps	grass + forb	167.88	173.27	175.290	6.568	4.906	15.333	0.029	1.073
CF_04	Cylindroiulus fulviceps	grass + forb	112.61	115.11	117.940	10.534	7.243	21.436	0.197	1.486
CF_05	Cylindroiulus fulviceps	grass + forb	115.87	118.84	121.840	53.284	39.577	63.515	0.629	4.092
CF_06	Cylindroiulus fulviceps	grass + forb	127.66	131.93	141.340	5.910	5.501	14.209	-0.017	0.999
CF_07	Cylindroiulus fulviceps	grass + forb	151.78	154.77	152.430	12.470	9.414	5.125	-0.297	0.592
CF_08	Cylindroiulus fulviceps	grass + forb	95.49	93.18	95.560	7.728	5.011	17.048	0.065	1.191
CF_09	Cylindroiulus fulviceps	grass + forb	159.05	159.16	166.500	7.848	6.346	14.466	-0.001	1.021
CF_10	Cylindroiulus fulviceps	grass + forb	145.35	141.87	143.390	5.682	4.867	10.158	-0.084	0.745
CF_01	Cylindroiulus fulviceps	shrub + grass + forb	135.81	145.49	148.800	6.288	4.605	3.102	-0.601	0.257
CF_02	Cylindroiulus fulviceps	shrub + grass + forb	124.42	126.75	125.820	5.968	4.194	13.111	-0.038	0.921
CF_03	Cylindroiulus fulviceps	shrub + grass + forb	167.88	173.27	175.290	6.278	4.560	14.223	-0.008	0.997
CF_04	Cylindroiulus fulviceps	shrub + grass + forb	112.61	115.11	117.940	6.326	5.759	16.911	0.085	1.182
CF_05	Cylindroiulus fulviceps	shrub + grass + forb	115.87	118.84	121.840	5.420	3.957	5.090	-0.390	0.409
CF_06	Cylindroiulus fulviceps	shrub + grass + forb	127.66	131.93	141.340	5.962	4.158	10.667	-0.155	0.755
CF_07	Cylindroiulus fulviceps	shrub + grass + forb	151.78	154.77	152.430	7.618	5.137	2.733	-0.543	0.323
CF_08	Cylindroiulus fulviceps	shrub + grass + forb	95.49	93.18	95.560	4.214	3.848	13.012	-0.067	0.914
CF_09	Cylindroiulus fulviceps	shrub + grass + forb	159.05	159.16	166.500	14.786	11.269	26.382	0.279	1.814
CF_10	Cylindroiulus fulviceps	shrub + grass + forb	145.35	141.87	143.390	5.324	4.703	9.807	-0.101	0.720