Citation:

j.marenvres.2013.10.010

Name: Negative effects of ocean acidification on two crustose coralline species using genetically homogeneous samples
Published: 2014-07-14
License: https://creativecommons.org/licenses/by/3.0/
Keywords: Benthos; Bottles or small containers/Aquaria (<20 L); Coast and continental shelf; Growth/Morphology; Hydrolithon samoense; Laboratory experiment; Lithophyllum kotschyanum; Macroalgae; North Pacific; Plantae; Rhodophyta; Single species; Temperate

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Abstract:

We evaluated acidification effects on two crustose coralline algal species common to Pacific coral reefs, Lithophyllum kotschyanum and Hydrolithon samoense. We used genetically homogeneous samples of both species to eliminate misidentification of species. The growth rates and percent calcification of the walls of the epithallial cells (thallus surface cells) of both species decreased with increasing pCO2. However, elevated pCO2 more strongly inhibited the growth of L. kotschyanum versus H. samoense. The trend of decreasing percent calcification of the cell wall did not differ between these species, although intercellular calcification of the epithallial cells in L. kotschyanum was apparently reduced at elevated pCO2, a result that might indicate that there are differences in the solubility or density of the calcite skeletons of these two species. These results can provide knowledge fundamental to future studies of the physiological and genetic mechanisms that underlie the response of crustose coralline algae to environmental stresses.

Keyword(s):

Benthos; Bottles or small containers/Aquaria (<20 L); Coast and continental shelf; Growth/Morphology; Hydrolithon samoense; Laboratory experiment; Lithophyllum kotschyanum; Macroalgae; North Pacific; Plantae; Rhodophyta; Single species; Temperate

Further details:

Lavigne, Héloïse; Epitalon, Jean-Marie; Gattuso, Jean-Pierre (2014): seacarb: seawater carbonate chemistry with R. R package version 3.0. https://cran.r-project.org/package=seacarb

Project(s):

Ocean Acidification International Coordination Centre (OA-ICC)

Comment:

In order to allow full comparability with other ocean acidification data sets, the R package seacarb (Lavigne et al, 2014) was used to compute a complete and consistent set of carbonate system variables, as described by Nisumaa et al. (2010). In this dataset the original values were archived in addition with the recalculated parameters (see related PI). The date of carbonate chemistry calculation is 2014-07-11.

Parameter(s):

#	Name	Short Name	Unit	Principal Investigator	Method/Device	Comment
1	Species	Species		Kato, A
2	Partial pressure of carbon dioxide (water) at sea surface temperature (wet air)	pCO2water_SST_wet	µatm	Kato, A		treatment
3	Aquarium number	Aquarium		Kato, A
4	Identification	ID		Kato, A		sample
5	Mass change	Mass chn	g	Kato, A		for 8 weeks
6	Number of points	Points	#	Kato, A		on cell walls
7	Number of points	Points	#	Kato, A		not on cell walls
8	Salinity	Sal		Kato, A
9	Temperature, water	Temp	°C	Kato, A
10	Temperature, water, standard deviation	Temp std dev	±	Kato, A
11	pH	pH		Kato, A	Potentiometric	total scale
12	pH, standard deviation	pH std dev	±	Kato, A	Potentiometric	total scale
13	Alkalinity, total	AT	µmol/kg	Kato, A	Potentiometric titration
14	Alkalinity, total, standard deviation	AT std dev	±	Kato, A	Potentiometric titration
15	Partial pressure of carbon dioxide (water) at sea surface temperature (wet air)	pCO2water_SST_wet	µatm	Kato, A	Calculated using CO2calc
16	Partial pressure of carbon dioxide, standard deviation	pCO2 std dev	±	Kato, A	Calculated using CO2calc
17	Bicarbonate ion	[HCO3]-	µmol/kg	Kato, A	Calculated using CO2calc
18	Bicarbonate ion, standard deviation	[HCO3]- std dev	±	Kato, A	Calculated using CO2calc
19	Carbonate ion	[CO3]2-	µmol/kg	Kato, A	Calculated using CO2calc
20	Carbonate ion, standard deviation	[CO3]2- std dev	±	Kato, A	Calculated using CO2calc
21	Carbon dioxide	CO2	µmol/kg	Kato, A	Calculated using CO2calc
22	Carbon dioxide, standard deviation	CO2 std dev	±	Kato, A	Calculated using CO2calc
23	Calcite saturation state	Omega Cal		Kato, A	Calculated using CO2calc
24	Calcite saturation state, standard deviation	Omega Cal std dev	±	Kato, A	Calculated using CO2calc
25	Aragonite saturation state	Omega Arg		Kato, A	Calculated using CO2calc
26	Aragonite saturation state, standard deviation	Omega Arg std dev	±	Kato, A	Calculated using CO2calc
27	Carbonate system computation flag	CSC flag		Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
28	Carbon dioxide	CO2	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
29	Partial pressure of carbon dioxide (water) at sea surface temperature (wet air)	pCO2water_SST_wet	µatm	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
30	Fugacity of carbon dioxide (water) at sea surface temperature (wet air)	fCO2water_SST_wet	µatm	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
31	Bicarbonate ion	[HCO3]-	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
32	Carbonate ion	[CO3]2-	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
33	Carbon, inorganic, dissolved	DIC	µmol/kg	Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
34	Aragonite saturation state	Omega Arg		Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)
35	Calcite saturation state	Omega Cal		Yang, Yan	Calculated using seacarb after Nisumaa et al. (2010)

License:

Creative Commons Attribution 3.0 Unported (CC-BY-3.0)

Status:

Curation Level: Enhanced curation (CurationLevelC)

Size:

4980 data points

Data

Download dataset as tab-delimited text — use the following character encoding:

1 Species	2 pCO2water_SST_wet [µatm] (treatment)	3 Aquarium	4 ID (sample)	5 Mass chn [g] (for 8 weeks)	6 Points [#] (on cell walls)	7 Points [#] (not on cell walls)	8 Sal	9 Temp [°C]	10 Temp std dev [±]	11 pH (total scale, Potentiometric)	12 pH std dev [±] (total scale, Potentiometric)	13 AT [µmol/kg] (Potentiometric titration)	14 AT std dev [±] (Potentiometric titration)	15 pCO2water_SST_wet [µatm] (Calculated using CO2calc)	16 pCO2 std dev [±] (Calculated using CO2calc)	17 [HCO3]- [µmol/kg] (Calculated using CO2calc)	18 [HCO3]- std dev [±] (Calculated using CO2calc)	19 [CO3]2- [µmol/kg] (Calculated using CO2calc)	20 [CO3]2- std dev [±] (Calculated using CO2calc)	21 CO2 [µmol/kg] (Calculated using CO2calc)	22 CO2 std dev [±] (Calculated using CO2calc)	23 Omega Cal (Calculated using CO2calc)	24 Omega Cal std dev [±] (Calculated using CO2calc)	25 Omega Arg (Calculated using CO2calc)	26 Omega Arg std dev [±] (Calculated using CO2calc)	27 CSC flag (Calculated using seacarb afte...)	28 CO2 [µmol/kg] (Calculated using seacarb afte...)	29 pCO2water_SST_wet [µatm] (Calculated using seacarb afte...)	30 fCO2water_SST_wet [µatm] (Calculated using seacarb afte...)	31 [HCO3]- [µmol/kg] (Calculated using seacarb afte...)	32 [CO3]2- [µmol/kg] (Calculated using seacarb afte...)	33 DIC [µmol/kg] (Calculated using seacarb afte...)	34 Omega Arg (Calculated using seacarb afte...)	35 Omega Cal (Calculated using seacarb afte...)
Lithophyllum kotschyanum (macroalga)	1019	1A	1	0.06452			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	2	0.05993			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	3	0.06468			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	4	0.06298			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	5	0.05247			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	6	0.06545			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	7	0.07606			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	8	0.05988			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	9	0.03860			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1A	10	0.05491			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	51	0.07644			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	52	0.08786			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	53	0.08158			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	54	0.07496			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	55	0.08518			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	56	0.04979			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	57	0.07341			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	58	0.08134			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	59	0.09683			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	1B	60	0.06650			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	281	4A	11	0.05080			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	12	0.06155			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	13	0.05903			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	14	0.06158			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	15	0.05957			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	16	0.06024			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	17	0.06243			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	18	0.05226			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	19	0.04739			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4A	20	0.06364			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	41	0.05981			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	42	0.05301			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	43	0.06766			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	44	0.07200			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	45	0.06886			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	46	0.07847			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	47	0.05301			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	48	0.08288			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	49	0.08296			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	4B	50	0.07788			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	418	5A	21	0.06250			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	22	0.06905			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	23	0.07059			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	24	0.04905			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	25	0.05677			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	26	0.03839			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	27	0.06254			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	28	0.05279			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	29	0.06418			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5A	30	0.06252			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	31	0.07864			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	32	0.06512			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	33	0.07848			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	34	0.07333			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	35	0.05226			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	36	0.06081			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	37	0.06980			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	38	0.06558			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	39	0.05637			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	5B	40	0.07506			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	1019	1A	1	0.00146			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	7	0.00214			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	13	-0.00010			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	19	0.00426			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	25	-0.00021			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	31	-0.00011			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	37	0.00429			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	43	0.00105			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	49	-0.00147			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1A	55	0.00171			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	4	0.00025			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	10	0.00252			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	16	0.00180			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	22	0.00344			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	28	0.00142			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	34	0.00224			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	40	0.00275			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	46	0.00245			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	52	0.00300			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	1B	58	0.00189			34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	281	4A	2	0.00409			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	8	0.00738			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	14	0.00401			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	20	0.00179			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	26	0.00344			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	32	0.00463			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	38	0.01178			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	44	0.00010			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	50	0.00143			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4A	56	0.00090			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	5	0.00522			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	11	0.00309			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	17	0.00516			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	23	0.00158			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	29	0.00648			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	35	0.00533			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	41	0.00437			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	47	0.00249			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	53	0.00333			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	4B	59	0.00098			34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	418	5A	3	0.00152			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	9	0.00169			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	15	0.00149			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	21	0.00228			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	27	0.00185			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	33	0.00075			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	39	0.00260			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	45	0.00228			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	51	0.00113			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5A	57	0.00150			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	6	0.00938			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	12	0.00414			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	18	0.00485			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	24	0.00491			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	30	0.00251			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	36	0.00809			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	42	0.00350			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	48	0.00266			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	54	0.00476			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	5B	60	0.00432			34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	1019	B	1		31	19	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	B	2		31	19	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	B	3		35	15	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	B	4		32	18	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	1019	B	5		36	14	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Lithophyllum kotschyanum (macroalga)	281	A	6		38	12	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	B	7		43	7	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	B	8		33	17	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	B	9		32	18	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	281	B	10		39	11	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Lithophyllum kotschyanum (macroalga)	418	A	11		39	11	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	B	12		36	14	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	B	13		36	14	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	B	14		34	16	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Lithophyllum kotschyanum (macroalga)	418	B	15		30	20	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	1019	A	16		45	5	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	A	17		39	11	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	A	18		30	20	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	A	19		39	11	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	1019	A	20		38	12	34.1	27.2	0.2	7.71	0.06	2279	12	960	143	1983	35	121	16	26.0	4	2.9	0.4	2.0	0.3	8	26	980	977	1992	117	2136	1.89	2.85
Hydrolithon samoense (macroalga)	281	A	21		44	6	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	A	22		45	5	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	A	23		39	11	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	A	24		42	8	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	281	A	25		45	5	34.1	27.3	0.2	8.18	0.06	2279	12	263	45	1585	61	282	27	7.1	1	6.9	0.7	4.6	0.4	8	7	266	265	1596	278	1881	4.49	6.77
Hydrolithon samoense (macroalga)	418	A	26		41	9	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	A	27		37	13	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	A	28		37	13	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	A	29		38	12	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35
Hydrolithon samoense (macroalga)	418	A	30		44	6	34.1	27.3	0.1	8.04	0.03	2279	12	399	37	1737	34	221	16	11.0	1	5.4	0.4	3.6	0.3	8	11	401	400	1740	220	1971	3.55	5.35