Citation:

j.orggeochem.2013.03.005

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Abstract:

In groundwater-fed fen peatlands, the surface biomass decays rapidly and, as a result, highly humified peat is formed. A high degree of humification constrains palaeoecological studies because reliable identification of plant remains is hampered. Organic geochemistry techniques as a means of identifying historical plant communities have been successfully applied tobog peat. The method has also been applied to fen peat, but without reference to the composition of fen plants. We have applied selected organic geochemistry methods to determine the composition of the neutral lipid fractions from 12 living fen plants, to investigate the potential for the distributions to characterize and separate different fen plants and plant groups. Our results show correspondence with previous studies, e.g. C23 and C25n-alkanes dominating Sphagnum spp. and C27 to C31 alkanes dominating vascular plants. However, we also found similarities in n-alkane distributions between Sphagnum spp. and the below ground parts of some vascular plants. We tested the efficiency of different n-alkane ratios to separate species and plant groups. The ratios used for bog studies (e.g. n-C23/n-C25 and n-C23/n-C29) did not work as consistently for fen plants. Some differences in sterol distribution were found between vascular plants and mosses; in general vascular plants had a higher concentration of sterols. When distributions of n-alkanes, n-alkane ratios and sterols were all included as variables, redundancy analysis (RDA) separated different plant groups into their own clusters. Our results imply that the pattern for bog biomarkers cannot directly be applied to fen environments. Nevertheless, they encourage further testing to determine whether or not the identification of plant groups, plants or plant parts from highly humified peat is possible by applying fen species-specific biomarker proxies.

Coverage:

Latitude: 64.750000 * Longitude: 24.700000

Event(s):

Finn_plants * Latitude: 64.750000 * Longitude: 24.700000 * Location: Finland * Method/Device: Sampling by hand (HAND)

Parameter(s):

#	Name	Short Name	Unit	Principal Investigator	Comment
1	Species	Species		Ronkainen, Tiina M
2	Sample type	Samp type		Ronkainen, Tiina M
3	n-Alkane C17	C17	ng/g	Ronkainen, Tiina M
4	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C17
5	n-Alkane C18	C18	ng/g	Ronkainen, Tiina M
6	n-Alkane C19	C19	ng/g	Ronkainen, Tiina M
7	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C19
8	n-Alkane C20	C20	ng/g	Ronkainen, Tiina M
9	n-Alkane C21	C21	ng/g	Ronkainen, Tiina M
10	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C21
11	n-Alkane C22	C22	ng/g	Ronkainen, Tiina M
12	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C22
13	n-Alkane C23	C23	ng/g	Ronkainen, Tiina M
14	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C23
15	n-Alkane C24	C24	ng/g	Ronkainen, Tiina M
16	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C24
17	n-Alkane C25	C25	ng/g	Ronkainen, Tiina M
18	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C25
19	n-Alkane C26	C26	ng/g	Ronkainen, Tiina M
20	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C26
21	n-Alkane C27	C27	ng/g	Ronkainen, Tiina M
22	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C27
23	n-Alkane C28	C28	ng/g	Ronkainen, Tiina M
24	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C28
25	n-Alkane C29	C29	ng/g	Ronkainen, Tiina M
26	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C29
27	n-Alkane C30	C30	ng/g	Ronkainen, Tiina M
28	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C30
29	n-Alkane C31	C31	ng/g	Ronkainen, Tiina M
30	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C31
31	n-Alkane C33	C33	ng/g	Ronkainen, Tiina M
32	Standard error	Std e	±	Ronkainen, Tiina M	n-Alkane C33
33	n-Alkane C35	C35	ng/g	Ronkainen, Tiina M
34	Squalene	Squalene	µg/g	Ronkainen, Tiina M
35	Standard error	Std e	±	Ronkainen, Tiina M	Squaline
36	n-Alkane, C21/C23 ratio	C21/C23		Ronkainen, Tiina M
37	Standard error	Std e	±	Ronkainen, Tiina M	n-C21/n-C23
38	n-Alkane, C23/C21 ratio	C23/C21		Ronkainen, Tiina M
39	Standard error	Std e	±	Ronkainen, Tiina M	n-C23/n-C21
40	n-Alkane, C23/C25 ratio	C23/C25		Ronkainen, Tiina M
41	Standard error	Std e	±	Ronkainen, Tiina M	n-C23/n-C25
42	n-Alkane, C23/C27 ratio	C23/C27		Ronkainen, Tiina M
43	Standard error	Std e	±	Ronkainen, Tiina M	n-C23/n-C27
44	n-Alkane, C23/C29 ratio	C23/C29		Ronkainen, Tiina M
45	Standard error	Std e	±	Ronkainen, Tiina M	n-C23/n-C29
46	n-Alkane, C23/C31 ratio	C23/C31		Ronkainen, Tiina M
47	Standard error	Std e	±	Ronkainen, Tiina M	n-C23/n-C31
48	n-Alkane, C25/C21 ratio	C25/C21		Ronkainen, Tiina M
49	Standard error	Std e	±	Ronkainen, Tiina M	n-C25/n-C21
50	n-Alkane, C25/C23 ratio	C25/C23		Ronkainen, Tiina M
51	Standard error	Std e	±	Ronkainen, Tiina M	n-C25/n-C23
52	n-Alkane, C25/C29 ratio	C25/C29		Ronkainen, Tiina M
53	Standard error	Std e	±	Ronkainen, Tiina M	n-C25/n-C29
54	n-Alkane, C31/C27 ratio	C31/C27		Ronkainen, Tiina M
55	Standard error	Std e	±	Ronkainen, Tiina M	n-C31/n-C27
56	n-Alkane, C31/C29 ratio	C31/C29		Ronkainen, Tiina M
57	Standard error	Std e	±	Ronkainen, Tiina M	n-C31/n-C29
58	n-Alkane, C33/C31 ratio	C33/C31		Ronkainen, Tiina M
59	Standard error	Std e	±	Ronkainen, Tiina M	n-C33/n-C31
60	Ratio	Ratio		Ronkainen, Tiina M	Paq
61	Standard error	Std e	±	Ronkainen, Tiina M	Paq
62	Ratio	Ratio		Ronkainen, Tiina M	Pwax
63	Standard error	Std e	±	Ronkainen, Tiina M	Pwax
64	n-Alkane, C23/(C23+C29) ratio	C23/(C23+C29)		Ronkainen, Tiina M
65	Standard error	Std e	±	Ronkainen, Tiina M	n-C23/(n-C23+n-C29)
66	n-Alkane, C25/(C25+C29) ratio	C25/(C25+C29)		Ronkainen, Tiina M
67	Standard error	Std e	±	Ronkainen, Tiina M	n-C25/(n-C25+n-C29)
68	n-Alkane, C23/(C27+C31) ratio	C23/(C27+C31)		Ronkainen, Tiina M
69	Standard error	Std e	±	Ronkainen, Tiina M	n-C23/(n-C27+n-C31)
70	Ratio	Ratio		Ronkainen, Tiina M	ACLn-C17-n-C35
71	Standard error	Std e	±	Ronkainen, Tiina M	ACLn-C17-n-C35
72	n-Alcohol C12	n-C12-ol	µg/g	Ronkainen, Tiina M
73	Standard error	Std e	±	Ronkainen, Tiina M	n-C12-ol
74	n-Alcohol C13	n-C13-ol	µg/g	Ronkainen, Tiina M
75	n-Alcohol C14	n-C14-ol	µg/g	Ronkainen, Tiina M
76	Standard error	Std e	±	Ronkainen, Tiina M	n-C14-ol
77	n-Alcohol C15	n-C15-ol	µg/g	Ronkainen, Tiina M
78	Standard error	Std e	±	Ronkainen, Tiina M	n-C15-ol
79	n-Alcohol C16	n-C16-ol	µg/g	Ronkainen, Tiina M
80	Standard error	Std e	±	Ronkainen, Tiina M	n-C16-ol
81	n-Alcohol C17	n-C17-ol	µg/g	Ronkainen, Tiina M
82	Standard error	Std e	±	Ronkainen, Tiina M	n-C17-ol
83	n-Alcohol C18	n-C18-ol	µg/g	Ronkainen, Tiina M
84	Standard error	Std e	±	Ronkainen, Tiina M	n-C18-ol
85	n-Alcohol C19	n-C19-ol	µg/g	Ronkainen, Tiina M
86	Standard error	Std e	±	Ronkainen, Tiina M	n-C19-ol
87	n-Alcohol C20	n-C20-ol	µg/g	Ronkainen, Tiina M
88	Standard error	Std e	±	Ronkainen, Tiina M	n-C20-ol
89	n-Alcohol C21	n-C21-ol	µg/g	Ronkainen, Tiina M
90	Standard error	Std e	±	Ronkainen, Tiina M	n-C21-ol
91	n-Alcohol C22	n-C22-ol	µg/g	Ronkainen, Tiina M
92	Standard error	Std e	±	Ronkainen, Tiina M	n-C22-ol
93	n-Alcohol C23	n-C23-ol	µg/g	Ronkainen, Tiina M
94	Standard error	Std e	±	Ronkainen, Tiina M	n-C23-ol
95	n-Alcohol C24	n-C24-ol	µg/g	Ronkainen, Tiina M
96	Standard error	Std e	±	Ronkainen, Tiina M	n-C24-ol
97	n-Alcohol C25	n-C25-ol	µg/g	Ronkainen, Tiina M
98	Standard error	Std e	±	Ronkainen, Tiina M	n-C25-ol
99	n-Alcohol C26	n-C26-ol	µg/g	Ronkainen, Tiina M
100	Standard error	Std e	±	Ronkainen, Tiina M	n-C26-ol
101	n-Alcohol C27	n-C27-ol	µg/g	Ronkainen, Tiina M
102	Standard error	Std e	±	Ronkainen, Tiina M	n-C27-ol
103	n-Alcohol C28	n-C28-ol	µg/g	Ronkainen, Tiina M
104	Standard error	Std e	±	Ronkainen, Tiina M	n-C28-ol
105	n-Alcohol C29	n-C29-ol	µg/g	Ronkainen, Tiina M
106	Standard error	Std e	±	Ronkainen, Tiina M	n-C29-ol
107	n-Alcohol C30	n-C30-ol	µg/g	Ronkainen, Tiina M
108	Standard error	Std e	±	Ronkainen, Tiina M	n-C30-ol
109	n-Alcohol C32	n-C32-ol	µg/g	Ronkainen, Tiina M
110	Standard error	Std e	±	Ronkainen, Tiina M	n-C32-ol
111	n-Alcohol C34	n-C34-ol	µg/g	Ronkainen, Tiina M
112	Standard error	Std e	±	Ronkainen, Tiina M	n-C34-ol
113	n-Alcohol C35	n-C35-ol	µg/g	Ronkainen, Tiina M
114	Standard error	Std e	±	Ronkainen, Tiina M	n-C35-ol
115	Phytol	Phytol	µg/g	Ronkainen, Tiina M
116	Standard error	Std e	±	Ronkainen, Tiina M	Phytol
117	delta-Tocopherol	d-Tocopherol	µg/g	Ronkainen, Tiina M
118	Standard error	Std e	±	Ronkainen, Tiina M	Tocopherol-d
119	gamma-Tocopherol	g-Tocopherol	µg/g	Ronkainen, Tiina M
120	Cholesterol	Cholesterol	µg/g	Ronkainen, Tiina M
121	Standard error	Std e	±	Ronkainen, Tiina M	Cholesterol
122	24-Methylcholesta-5,22E-dien-3beta-ol	Brassicasterol	µg/g	Ronkainen, Tiina M
123	Standard error	Std e	±	Ronkainen, Tiina M	Brassicasterol
124	24-Methylcholest-5-en-3beta-ol	Capesterol	µg/g	Ronkainen, Tiina M
125	Standard error	Std e	±	Ronkainen, Tiina M	Campesterol
126	Sterols	Sterol	µg/g	Ronkainen, Tiina M	SterolA
127	Triterpenoids, plant-derived	Triterp plant	µg/g	Ronkainen, Tiina M	Triterpenoid-1
128	Stigmasterol	Stigmasterol	µg/g	Ronkainen, Tiina M
129	Standard error	Std e	±	Ronkainen, Tiina M	Stigmasterol
130	Ergost-8,24(28)-dien-3b-ol	Ergost-8,24(28)-dien-3b-ol	µg/g	Ronkainen, Tiina M
131	24-Methylcholest-7-en-3b-ol	24-Methylcholest-7-en-3b-ol	µg/g	Ronkainen, Tiina M
132	Agnosterol	Agnosterol	µg/g	Ronkainen, Tiina M
133	Sterols	Sterol	µg/g	Ronkainen, Tiina M	SterolB
134	Standard error	Std e	±	Ronkainen, Tiina M	SterolB
135	Triterpenol	Triterpenol	µg/g	Ronkainen, Tiina M
136	beta-Amyrin	b-Amyrin	µg/g	Ronkainen, Tiina M
137	Standard error	Std e	±	Ronkainen, Tiina M	beta-Amyrin
138	Obtusifoliol	Obtusifoliol	µg/g	Ronkainen, Tiina M
139	Standard error	Std e	±	Ronkainen, Tiina M	Obtusifoliol
140	24-Ethylcholest-5-en-3beta-ol	b-Sitosterol	µg/g	Ronkainen, Tiina M
141	Standard error	Std e	±	Ronkainen, Tiina M	beta-sitosterol
142	3-Stimastanol	3-Stimastanol	µg/g	Ronkainen, Tiina M
143	Standard error	Std e	±	Ronkainen, Tiina M	3-stimastanol
144	Stigmasta-3,5-dien-7-one	Stigmasta-3,5-dien-7-one	µg/g	Ronkainen, Tiina M
145	Sterols	Sterol	µg/g	Ronkainen, Tiina M	SterolC
146	Standard error	Std e	±	Ronkainen, Tiina M	SterolC
147	Lupeol	Lupeol	µg/g	Ronkainen, Tiina M
148	Standard error	Std e	±	Ronkainen, Tiina M	Lupeol
149	Gramisterol	Gramisterol	µg/g	Ronkainen, Tiina M
150	Standard error	Std e	±	Ronkainen, Tiina M	Gramisterol
151	Lanosterol	Lanosterol	µg/g	Ronkainen, Tiina M
152	Cycloartenol	Cycloartenol	µg/g	Ronkainen, Tiina M
153	Standard error	Std e	±	Ronkainen, Tiina M	Cycloartenol
154	Schottenol	Schottenol	µg/g	Ronkainen, Tiina M
155	Standard error	Std e	±	Ronkainen, Tiina M	Schottenol
156	Avenasterol	Avenasterol	µg/g	Ronkainen, Tiina M
157	Standard error	Std e	±	Ronkainen, Tiina M	Avenasterol
158	Sterols	Sterol	µg/g	Ronkainen, Tiina M	branched/cyclic alcohol
159	24-Methylene-cycloartenol	24-Methylene-cycloartenol	µg/g	Ronkainen, Tiina M
160	Standard error	Std e	±	Ronkainen, Tiina M	24-Methylenecycloartenol
161	Sterols	Sterol	µg/g	Ronkainen, Tiina M	SterolD
162	Standard error	Std e	±	Ronkainen, Tiina M	SterolD

License:

Creative Commons Attribution 3.0 Unported (CC-BY-3.0)

Size:

2596 data points

Data

Download dataset as tab-delimited text — use the following character encoding:

1 Species	2 Samp type	3 C17 [ng/g]	4 Std e [±] (n-Alkane C17)	5 C18 [ng/g]	6 C19 [ng/g]	7 Std e [±] (n-Alkane C19)	8 C20 [ng/g]	9 C21 [ng/g]	10 Std e [±] (n-Alkane C21)	11 C22 [ng/g]	12 Std e [±] (n-Alkane C22)	13 C23 [ng/g]	14 Std e [±] (n-Alkane C23)	15 C24 [ng/g]	16 Std e [±] (n-Alkane C24)	17 C25 [ng/g]	18 Std e [±] (n-Alkane C25)	19 C26 [ng/g]	20 Std e [±] (n-Alkane C26)	21 C27 [ng/g]	22 Std e [±] (n-Alkane C27)	23 C28 [ng/g]	24 Std e [±] (n-Alkane C28)	25 C29 [ng/g]	26 Std e [±] (n-Alkane C29)	27 C30 [ng/g]	28 Std e [±] (n-Alkane C30)	29 C31 [ng/g]	30 Std e [±] (n-Alkane C31)	31 C33 [ng/g]	32 Std e [±] (n-Alkane C33)	33 C35 [ng/g]	34 Squalene [µg/g]	35 Std e [±] (Squaline)	36 C21/C23	37 Std e [±] (n-C21/n-C23)	38 C23/C21	39 Std e [±] (n-C23/n-C21)	40 C23/C25	41 Std e [±] (n-C23/n-C25)	42 C23/C27	43 Std e [±] (n-C23/n-C27)	44 C23/C29	45 Std e [±] (n-C23/n-C29)	46 C23/C31	47 Std e [±] (n-C23/n-C31)	48 C25/C21	49 Std e [±] (n-C25/n-C21)	50 C25/C23	51 Std e [±] (n-C25/n-C23)	52 C25/C29	53 Std e [±] (n-C25/n-C29)	54 C31/C27	55 Std e [±] (n-C31/n-C27)	56 C31/C29	57 Std e [±] (n-C31/n-C29)	58 C33/C31	59 Std e [±] (n-C33/n-C31)	60 Ratio (Paq)	61 Std e [±] (Paq)	62 Ratio (Pwax)	63 Std e [±] (Pwax)	64 C23/(C23+C29)	65 Std e [±] (n-C23/(n-C23+n-C29))	66 C25/(C25+C29)	67 Std e [±] (n-C25/(n-C25+n-C29))	68 C23/(C27+C31)	69 Std e [±] (n-C23/(n-C27+n-C31))	70 Ratio (ACLn-C17-n-C35)	71 Std e [±] (ACLn-C17-n-C35)	72 n-C12-ol [µg/g]	73 Std e [±] (n-C12-ol)	74 n-C13-ol [µg/g]	75 n-C14-ol [µg/g]	76 Std e [±] (n-C14-ol)	77 n-C15-ol [µg/g]	78 Std e [±] (n-C15-ol)	79 n-C16-ol [µg/g]	80 Std e [±] (n-C16-ol)	81 n-C17-ol [µg/g]	82 Std e [±] (n-C17-ol)	83 n-C18-ol [µg/g]	84 Std e [±] (n-C18-ol)	85 n-C19-ol [µg/g]	86 Std e [±] (n-C19-ol)	87 n-C20-ol [µg/g]	88 Std e [±] (n-C20-ol)	89 n-C21-ol [µg/g]	90 Std e [±] (n-C21-ol)	91 n-C22-ol [µg/g]	92 Std e [±] (n-C22-ol)	93 n-C23-ol [µg/g]	94 Std e [±] (n-C23-ol)	95 n-C24-ol [µg/g]	96 Std e [±] (n-C24-ol)	97 n-C25-ol [µg/g]	98 Std e [±] (n-C25-ol)	99 n-C26-ol [µg/g]	100 Std e [±] (n-C26-ol)	101 n-C27-ol [µg/g]	102 Std e [±] (n-C27-ol)	103 n-C28-ol [µg/g]	104 Std e [±] (n-C28-ol)	105 n-C29-ol [µg/g]	106 Std e [±] (n-C29-ol)	107 n-C30-ol [µg/g]	108 Std e [±] (n-C30-ol)	109 n-C32-ol [µg/g]	110 Std e [±] (n-C32-ol)	111 n-C34-ol [µg/g]	112 Std e [±] (n-C34-ol)	113 n-C35-ol [µg/g]	114 Std e [±] (n-C35-ol)	115 Phytol [µg/g]	116 Std e [±] (Phytol)	117 d-Tocopherol [µg/g]	118 Std e [±] (Tocopherol-d)	119 g-Tocopherol [µg/g]	120 Cholesterol [µg/g]	121 Std e [±] (Cholesterol)	122 Brassicasterol [µg/g]	123 Std e [±] (Brassicasterol)	124 Capesterol [µg/g]	125 Std e [±] (Campesterol)	126 Sterol [µg/g] (SterolA)	127 Triterp plant [µg/g] (Triterpenoid-1)	128 Stigmasterol [µg/g]	129 Std e [±] (Stigmasterol)	130 Ergost-8,24(28)-dien-3b-ol [µg/g]	131 24-Methylcholest-7-en-3b-ol [µg/g]	132 Agnosterol [µg/g]	133 Sterol [µg/g] (SterolB)	134 Std e [±] (SterolB)	135 Triterpenol [µg/g]	136 b-Amyrin [µg/g]	137 Std e [±] (beta-Amyrin)	138 Obtusifoliol [µg/g]	139 Std e [±] (Obtusifoliol)	140 b-Sitosterol [µg/g]	141 Std e [±] (beta-sitosterol)	142 3-Stimastanol [µg/g]	143 Std e [±] (3-stimastanol)	144 Stigmasta-3,5-dien-7-one [µg/g]	145 Sterol [µg/g] (SterolC)	146 Std e [±] (SterolC)	147 Lupeol [µg/g]	148 Std e [±] (Lupeol)	149 Gramisterol [µg/g]	150 Std e [±] (Gramisterol)	151 Lanosterol [µg/g]	152 Cycloartenol [µg/g]	153 Std e [±] (Cycloartenol)	154 Schottenol [µg/g]	155 Std e [±] (Schottenol)	156 Avenasterol [µg/g]	157 Std e [±] (Avenasterol)	158 Sterol [µg/g] (branched/cyclic alcohol)	159 24-Methylene-cycloartenol [µg/g]	160 Std e [±] (24-Methylenecycloartenol)	161 Sterol [µg/g] (SterolD)	162 Std e [±] (SterolD)
Sphagnum fimbriatum		377		1643	0		0	4685		1563		21303		1760		36529		1121		14436		0		2294		0		675		0		0	0.0		0.2		4.5		0.6		1.5		9.3		31.6		7.8		1.7		15.9		0.0		0.3		0.0		1.0		0.2		0.9		0.9		1.4		24.7		0.0		0.0	21.4		0.0		86.2		7.5		39.4		3.2		16.1		13.0		301.6		44.3		471.1		27.3		229.5		7.1		56.4		0.0		8.6		0.0		0.0		0.0		400.3		0.0		0.0	47.6		29.2		574.6		0.0	0.0	402.1		6.3	4.6	1.3	0.0		0.0	0.0		42.9		550.7		0.0		13.6	0.0		0.0		18.4		0.0	8.1		0.0		0.0		0.0	21.5		0.0
Sphagnum subsecundum		1034		4528	519		0	13382		2434		23622		1677		13851		744		5120		277		2167		153		915		0		0	0.0		0.6		1.8		1.7		4.6		10.9		25.8		1.0		0.6		6.4		0.2		0.4		0.0		0.9		0.2		0.9		0.9		3.9		23.6		0.0		0.0	12.4		0.0		20.6		8.3		15.5		1.4		13.6		9.0		133.0		28.2		274.2		28.0		331.8		10.4		235.4		0.0		10.2		0.0		0.0		0.0		413.9		0.0		0.0	32.7		25.3		167.9		0.0	0.0	249.4		17.2	11.6	7.5	0.0		0.0	0.0		277.9		344.0		0.0		37.0	0.0		0.0		57.8		0.0	186.6		0.0		6.7		0.0	122.2		30.8
Sphagnum papillosum		0		0	0		0	5285	1869	1246		32639	3572	1179		11252	3160	486		1917	821	0		1147		260		1557	559	997	417	0	0.0		0.2	0.0	28.7	13.5	3.2	0.5	21.8	6.7	34.4		28.7	13.5	2.2	0.2	0.3	0.1	15.3		1.4	0.9	0.8		0.6	0.0	1.0	0.0	0.1	0.0	1.0	0.0	1.0	0.0		0.7	23.6	0.2	2.7		0.0	7.6	1.3	0.0		6.2	0.8	0.5		19.7	2.1	0.0		4.5	0.3	9.8	3.5	109.1	31.5	13.1	4.4	469.2	162.7	22.8	8.1	324.7	134.4	0.0		54.3	42.1	0.0		0.0		0.0		0.0		0.0	0.0	615.0	292.4	0.0		0.0	11.8	4.3	15.1	9.6	288.6	144.5	0.0	0.0	230.5	123.5	14.2	22.0	28.5	5.6	0.3	0.0	0.0		338.0	267.4	720.8	317.3	0.0		28.1	3.8	0.8	0.0		60.9	47.6	0.0	48.8		0.0		0.0		0.0	90.0	64.2	0.0
Sphagnum riparium		162		158	370		0	8383		627		12357		817		7648		302		2023		115		696		0		466		149		149	2.7		0.7		1.5		1.6		6.1		17.8		26.5		0.9		0.6		11.0		0.2		0.7		0.3		0.9		0.1		0.9		0.9		5.0		23.4		0.0		0.0	15.9		0.0		6.4		0.0		16.2		0.0		7.8		2.5		54.8		7.1		131.2		10.1		144.0		3.0		30.3		0.0		0.0		0.0		0.0		0.0		835.5		0.0		0.0	27.5		206.8		436.1		0.0	0.0	634.4		17.0	12.7	0.0	0.0		0.0	0.0		222.4		163.4		0.0		0.0	0.0		0.0		25.2		0.0	62.6		0.0		3.8		0.0	32.1		7.6
Warnstorfia exannulata		2342	398	0	0		0	608		520		1307	423	549		1491	432	408		2278	947	0		2178	205	583		0		978	399	0	0.0		0.2	0.1			0.9	0.0	0.3	0.1	0.6	0.1			3.3		1.2	0.0	0.7	0.1	0.0	0.0	0.0	0.0			0.5	0.1	0.8	0.1	0.4	0.1	0.4	0.0	0.3	0.1	23.2	1.2	0.0		0.0	86.6	44.9	9.3	5.4	46.9	26.3	9.4	6.5	106.6	53.1	8.5	5.3	40.5	20.2	3.9	1.4	67.2	32.7	4.8	1.7	181.5	73.9	12.0		495.1	196.8	8.2	4.9	839.1	299.4	0.0		63.9	21.5	0.0		0.0		0.0		2035.9	913.3	0.0		0.0	141.3	53.2	25.4	11.2	513.3	215.8	0.0	0.0	327.3	133.3	28.2	31.8	0.0	0.0		0.0	0.0		0.0		599.5	248.4	0.0		0.0	18.3	7.3	0.0		11.7		0.0	53.9		0.0		16.2		0.0	30.7		0.0
Warnstorfia exannulata rep		1057		157	173		176	435		464		1771		625		10662		837		10664		252		1915		0		4177		1880		0	0.0		0.2		4.1		0.2		0.2		0.9		0.4		24.5		6.0		5.6		0.4		2.2		0.5		0.7		0.6		0.5		0.8		0.1		26.7		0.0		0.0	86.5		0.0		62.6		117.5		1210.7		0.0		74198.9		221.3		3657.7		1503.3		36.5		1893.7		220.2		13301.0		302.6		0.0		0.0		0.0		0.0		0.0		437.9		0.0		0.0	0.0		10682.7		790.0		0.0	0.0	22444.1		299.9	14813.8	0.0	0.0		0.0	0.0		0.0		425.9		20978.1		0.0	0.0		0.0		0.0		0.0	1311.4		0.0		0.0		0.0	0.0
Carex livida	above-ground parts	0		0	352		0	1400		244		1038		506		1216		1481		7130		1079		7003		467		3630		144		144	4.6		1.3		0.7		0.9		0.1		0.1		0.3		0.9		1.2		0.2		0.5		0.5		0.0		0.2		0.9		0.1		0.1		0.1		27.5		0.0		0.0	3.4		0.0		52.7		67.3		31.7		2.1		16.0		2.4		22.9		2.4		79.6		3.9		116.5		18.5		1040.9		6.1		132.7		13.7		0.0		0.0		638.8		0.0		0.0	22.3		0.0		121.8		0.0	0.0	169.4		0.0	0.0	0.0	0.0		0.0	10.4		8.5		864.2		0.0		0.0	0.0		18.1		0.0		0.0	119.0		0.0		0.0		10.3	52.5		11.6
Carex livida	below-ground part	0		403	1505		0	5699		292		3000		0		480		0		744		0		599		0		639		0		0	4.3		1.9		0.5		6.3		4.0		5.0		4.7		0.1		0.2		0.8		0.9		1.1		0.0		0.7		0.4		0.8		0.4		2.2		22.6		0.0		0.0	13.2		0.0		189.2		6.3		307.5		1.6		46.8		0.0		27.0		0.0		66.6		3.1		95.5		12.5		626.2		0.0		94.7		18.3		0.0		0.0		7.2		0.0		0.0	72.7		35.4		494.9		27.8	0.0	699.8		0.0	0.0	0.0	0.0		0.0	0.0		13.4		2252.9		316.7		0.0	0.0		22.5		0.0		0.0	94.2		0.0		0.0		102.0	63.3		0.0
Carex livida rep	below-ground part	0		87	71		90	231		330		2508		891		1348		525		554		230		604		0		266		0		0	0.5		0.2		4.3		1.8		1.9		2.1		4.7		2.4		0.6		1.2		0.4		0.4		0.0		0.7		0.4		0.7		0.5		1.3		25.3		0.0		0.0	18.3		0.0		1663.0		23.7		977.1		0.0		34.5		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		34.6		0.0		0.0	60.1		465.4		543.3		0.0	0.0	0.0		0.0	0.0	0.0	0.0		0.0	0.0		0.0		2532.0		0.0		0.0	0.0		0.0		0.0		0.0	96.1		0.0		0.0		0.0	0.0		0.0
Carex nigra	above-ground parts	0		0	0		0	737		0		3020		592		1541		1421		12287		261		1836		0		0		0		0	3.5		0.2		4.1		2.0		0.2		1.6				2.1		0.5		0.8		0.0		0.0				0.7		0.8		0.6		0.5		0.2		26.2		0.0		0.0	0.0		0.0		50.0		359.6		32.2		0.0		70.7		13.0		125.3		10.0		517.5		0.0		251.3		111.1		25635.0		77.7		272.4		0.0		0.0		0.0		7510.9		0.0		0.0	100.9		0.0		1010.7		0.0	0.0	151.3		0.0	0.0	0.0	58.2		0.0	0.0		83.0		8575.7		517.3		0.0	0.0		253.3		0.0		0.0	472.5		0.0		70.3		0.0	193.0		95.9
Carex nigra	below-ground part	0		0	0		0	1140		0		1704		0		257		0		5191		0		2399		0		214		0		0	0.0		0.7		1.5		6.6		0.3		0.7		8.0		0.2		0.2		0.1		0.0		0.1		0.0		0.4		0.8		0.4		0.1		0.3		26.2		0.0		0.0	11.8		1.2		250.8		20.7		23.9		3.3		159.8		21.0		363.2		27.1		1875.7		13.1		386.7		17.7		1487.5		0.0		339.6		0.0		0.0		0.0		7.7		0.0		0.0	36.7		0.0		615.5		0.0	0.0	52.0		0.0	0.0	0.0	0.0		0.0	65.1		22.3		3849.6		414.7		0.0	0.0		231.1		0.0		0.0	108.2		0.0		15.6		0.0	101.9		24.7
Carex nigra rep	below-ground part	0		103	0		0	127		179		720		499		779		533		806		168		636		0		243		0		0	0.2		0.2		5.7		0.9		0.9		1.1		3.0		6.1		1.1		1.2		0.3		0.4		0.0		0.6		0.5		0.5		0.6		0.7		26.1		0.0		0.0	0.0		0.0		1230.8		0.0		28.8		0.0		85.3		13.6		103.6		6.0		105.7		0.0		0.0		0.0		37.7		0.0		0.0		0.0		0.0		0.0		11.8		0.0		0.0	0.0		24.9		797.7		0.0	0.0	79.4		0.0	0.0	0.0	0.0		0.0	218.5		0.0		8185.3		1017.0		0.0	0.0		59.7		0.0		0.0	0.0		0.0		0.0		0.0	0.0		0.0
Carex rostrata	above-ground parts	0		0	0		0	445		530		3859	1098	1258	290	15471	2491	2138	456	66595	6402	4443	804	188799	30847	1076	207	32247	6060	16178	2881	841	0.0		0.0	0.0	0.1	0.0	0.2	0.0	0.1	0.0	0.0	0.0	0.1	0.0	45.7		4.3	0.5	0.1	0.0	0.5	0.1	0.2	0.0	0.5	0.0	0.1	0.0	0.9	0.0	0.0	0.0	0.1	0.0	0.0	0.0	28.7	0.1	0.0		0.0	17.9	2.8	0.0		18.9	2.9	166.3		68.6	11.3	4.4	2.1	404.1	87.9	19.8	5.9	991.5	209.1	24.5	7.1	682.0	175.5	31.0	9.7	168.4	47.6	38.1	11.9	178.0	32.3	18.1	6.5	120.6	21.6	10.3	3.6	0.0		0.0		1502.3	1036.7	0.0		0.0	67.3	21.4	0.0		250.6	87.3	0.0	0.0	124.8	49.5	15.4	0.0	0.0	57.8		0.0	0.0		9.4	3.4	2525.2	772.6	151.0	69.2	0.0	14.8	1.5	0.0		0.0		0.0	50.4	30.1	0.0		17.0	11.4	0.0	42.8	26.6	19.6	14.0
Carex rostrata	below-ground part	0		0	6845	3588	520	11144	4739	563	152	24731	10196	1334	575	15472	6565	600		4848	2199	0		21850	7228	617	242	3520	1367	550	46	0	118.7	109.6	0.4	0.0	6.9	0.4	1.6	0.0	5.3	0.2	1.1	0.1	6.9	0.4	1.4	0.0	0.6	0.0	0.7	0.1	0.8	0.1	0.2	0.0	0.2	0.1	0.6	0.0	0.4	0.0	0.5	0.0	0.4	0.0	3.0	0.1	25.0	0.2	2.4	0.1	1.4	9.8	1.0	2.4	0.4	105.8	12.3	80.0	78.2	67.1	5.4	10.9	2.2	920.1	97.2	36.3	6.0	1011.0	118.2	34.3	7.4	1224.5	144.2	46.0	10.7	630.4	75.4	9.3	3.0	192.2	22.0	4.3		104.6	3.2	10.4	3.2	0.0		0.0		35.1	13.7	0.0		0.0	90.2	17.9	37.3	8.2	387.7	88.0	0.0	0.0	189.3	25.7	6.4	0.0	0.0	0.0		89.2	72.3	32.5	0.0		3662.1	592.1	248.2	47.9	0.0	55.4	6.6	28.5	10.6	0.0		0.0	54.2	19.7	0.0		7.5		0.0	19.4	6.9	10.0	4.8
Carex rostrata rep	below-ground part	0		0	0		0	201		147		987		227		739		146		275		0		506		0		0		0		0	1.5		0.2		4.9		1.3		3.6		2.0				3.7		0.7		1.5		0.0		0.0				0.8		0.3		0.7		0.6		3.6		24.9		0.0		0.0	2.4		0.0		190.8		0.0		24.6		0.0		0.0		134.2		114.0		0.0		92.1		0.0		25.8		0.0		0.0		0.0		0.0		0.0		0.0		0.0		7.6		0.0		0.0	60.0		39.5		901.5		0.0	0.0	303.0		0.0	0.0	0.0	0.0		0.0	0.0		0.0		9177.5		659.0		0.0	0.0		0.0		0.0		0.0	376.9		0.0		0.0		0.0	0.0		0.0
Carex lasiocarpa	above-ground parts	196		0	0		190	152		0		670		170		1487		184		8459		676		52792		447		9631		768		768	2.1		0.2				0.5		0.1		0.0		0.1		9.8		2.2		0.0		1.1		0.2		0.1		0.0		1.0		0.0		0.0		0.0		28.9		0.0		0.0	8.6		0.0		0.0		15.8		8.9		0.0		9.7		1.3		51.3		2.4		203.0		6.2		309.8		4.6		370.9		7.3		221.6		36.2		0.0		0.0		432.7		0.0		0.0	16.9		0.0		214.6		8.2	0.0	25.5		2.9	0.0	0.0	0.0		0.0	25.4		3.8		1172.0		85.9		0.0	0.0		0.0		0.0		0.0	176.2		0.0		0.0		0.0	15.9		4.9
Carex lasiocarpa	below-ground part	0		0	370		0	2204		986		15703		1444		3443		0		803		0		1701		0		1127		0		0	4.7		0.1		15.9		4.6		19.6		9.2		13.9		1.6		0.2		2.0		1.4		0.7		0.0		0.9		0.2		0.9		0.7		8.1		23.9		0.0		0.0	19.6		0.0		12.7		6.7		17.8		4.1		23.7		11.8		269.6		37.4		1093.9		36.5		859.5		9.9		277.9		0.0		118.6		43.3		0.0		0.0		15.9		0.0		0.0	78.4		11.7		694.9		24.7	0.0	73.0		0.0	0.0	0.0	0.0		72.9	0.0		22.9		3712.9		215.4		0.0	0.0		123.6		0.0		0.0	1296.9		0.0		0.0		0.0	43.3		13.8
Carex lasiocarpa rep	below-ground part	0		87	71		90	231		330		2508		891		1348		525		554		230		604		0		266		0		0	0.4		0.1		10.9		1.9		4.5		4.2		9.4		5.8		0.5		2.2		0.5		0.4		0.0		0.8		0.3		0.8		0.7		3.1		24.8		0.0		0.0	3.3		0.0		41.6		0.0		7.7		0.0		3.6		1.8		100.0		13.2		339.6		6.9		90.9		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0		0.0	67.5		0.0		806.2		0.0	0.0	54.7		0.0	0.0	0.0	0.0		0.0	0.0		0.0		5126.7		431.7		0.0	0.0		0.0		0.0		0.0	6982.9		0.0		0.0		0.0	0.0		0.0
Eriop.ang	above-ground parts	0		0	0		0	363		115		3172		519		6898		680		10208		282		4863		0		462		0		0	7.9		0.1		8.7		0.5		0.3		0.7		6.9		19.0		2.2		1.4		0.0		0.1		0.0		0.7		0.6		0.4		0.6		0.3		26.3		0.0		0.0	3.3		0.0		4.2		79.7		8.3		0.0		52.1		10.7		560.0		17.7		640.2		16.8		911.9		19.6		592.0		4.7		128.0		9.6		0.0		0.0		983.2		0.0		0.0	31.0		0.0		108.3		0.0	0.0	36.6		1.4	3.8	0.0	0.0		0.0	17.2		15.9		1433.5		0.0		0.0	0.0		22.6		0.0		0.0	215.4		0.0		7.1		0.0	47.0		14.0
Eriop.ang	below-ground part	0		0	0		0	2063		262		7293		280		9212		455		14529		0		5744		0		370		0		0	56.3		0.3		3.5		0.8		0.5		1.3		19.7		4.5		1.3		1.6		0.0		0.1		0.0		0.7		0.6		0.6		0.6		0.5		25.8		0.0		0.0	4.6		0.0		10.1		8.0		38.6		1.6		95.0		13.3		459.2		21.5		728.7		26.9		1184.8		44.7		952.3		0.0		239.9		18.5		0.0		38.8		26.9		0.0		0.0	60.8		0.0		375.5		0.0	0.0	522.8		8.6	13.7	0.0	0.0		0.0	0.0		8.8		3442.8		160.0		0.0	0.0		0.0		0.0		0.0	106.5		0.0		9.0		0.0	36.3		28.0
Eriop.ang rep	below-ground part	0		46	0		0	820		683		4027		3280		15318		3700		26736		1383		16740		167		952		0		0	0.4		0.2		4.9		0.3		0.2		0.2		4.2		18.7		3.8		0.9		0.0		0.1		0.0		0.5		0.7		0.2		0.5		0.1		26.8		0.0		0.0	0.0		0.0		0.0		0.0		23.8		2.7		130.6		17.8		892.9		15.1		682.2		11.9		713.4		16.4		315.7		0.0		0.0		0.0		0.0		0.0		3481.1		0.0		0.0	0.0		0.0		308.0		0.0	0.0	24.0		0.0	0.0	0.0	0.0		0.0	0.0		0.0		5374.7		341.0		0.0	0.0		0.0		49.7		0.0	638.6		0.0		0.0		0.0	0.0		0.0
Menyanthes trifoliata	above-ground parts	0		0	0		0	275	12	0		1228	114	171		1113	104	147		1174	88	0		1467	168	0		949		0		0	61.7	53.0	0.1	0.1	1.1		1.1	0.0	1.0	0.0	0.9	0.1	1.1		4.2	0.3	0.9	0.0	0.8	0.1	0.3	0.3	0.3	0.3	0.0		0.6	0.0	0.6	0.0	0.5	0.0	0.4	0.0	0.9	0.2	26.3	0.3	1.2		0.0	5.1	1.0	0.0		5.7	0.5	2.3		34.1	6.1	1.5		367.3	86.0	12.2	2.8	941.7	221.5	16.0	2.8	658.3	160.5	33.5	7.2	503.9	116.4	22.9	3.6	436.9	102.7	9.4	1.9	91.4	15.2	39.4	8.8	6.6		0.0		3030.5	818.9	44.1	41.8	20.5	15.6	5.4	0.0		47.0		0.0	17.0	1483.6	249.9	0.0	0.0	0.0	0.0		0.0	0.0		31.6		0.0		0.0		0.0	0.0		0.0		0.0		0.0	83.5	38.1	678.0	83.5	97.1	5.2	0.0	23.6	0.4	21.9	2.0
Menyanthes trifoliata	below-ground part	0		0	0		542	38042		10392		25548		3382		8184		490		1964		0		617		0		374		190		190	0.0		1.5		0.7		3.1		13.0		41.4		68.2		0.2		0.3		13.3		0.2		0.6		0.5		1.0		0.1		1.0		0.9		10.9		22.4		0.0		0.0	4.1		0.0		10.9		67.7		28.1		0.0		38.7		5.1		81.9		3.7		204.6		8.8		220.8		9.6		137.3		0.0		0.0		0.0		0.0		0.0		297.4		288.4		40.2	24.8		0.0		227.7		0.0	788.8	3647.4		0.0	0.0	0.0	0.0		0.0	0.0		0.0		0.0		0.0		0.0	0.0		293.6		0.0		718.0	443.9		1029.4		122.4		0.0	0.0		0.0
Menyanthes trifoliata rep	below-ground part	0		104	105		415	22793		5793		19103		3228		6091		796		2301		296		819		0		0		0		0	0.4		1.2		0.8		3.1		8.3		23.3				0.3		0.3		7.4		0.0		0.0				1.0		0.1		1.0		0.9		8.3		22.6		0.0		0.0	0.0		0.0		0.0		0.0		14.9		0.0		3.1		2.7		27.7		2.3		53.5		0.0		50.7		0.0		14.3		0.0		0.0		0.0		0.0		0.0		204.8		100.4		225.8	0.0		0.0		2145.6		0.0	0.0	639.7		0.0	0.0	0.0	0.0		0.0	116.4		0.0		0.0		0.0		0.0	0.0		1009.9		0.0		0.0	0.0		0.0		0.0		0.0	0.0		0.0
Potentilla palustris	above-ground parts	0		0	0		0	264		0		4570	717	595	18	8656	1659	769	487	37322	10049	454		69762	22288	1331	501	497565	182606	364996	133442	743	115.8	115.2	0.0	0.0	0.0	0.0	0.5	0.1	0.1	0.0	0.1	0.0	0.0	0.0	24.6		1.9	0.2	0.1	0.0	12.4	1.7	6.8	0.6	0.7	0.0	0.0	0.0	1.0	0.0	0.1	0.0	0.1	0.0	0.0	0.0	31.3	0.1	1.4	0.4	0.0	2.0	0.4	1.1		6.8	1.1	8.1	6.2	66.5	11.1	1.5	0.2	518.3	99.0	46.2	9.6	3045.2	686.6	47.7	9.8	1262.5	294.2	56.8	15.7	1142.7	333.6	88.9	30.0	742.4	258.4	21.4	7.1	91.7	32.8	53.7	20.3	19.8	8.7	0.0		1257.7	182.7	0.0		0.0	255.3	54.4	0.0		47.1	4.9	0.0	0.0	8.7	1.2	0.0	0.0	0.0	0.0		0.0	38.1		6.6		1945.6	196.4	164.4	8.4	0.0	32.2	3.3	0.0		0.0		129.7	40.0	7.6	0.0		19.9		0.0	27.5	6.8	59.3
Potentilla palustris	below-ground part	0		0	0		0	0		0		256		0		688		246		2813		145		7099		0		14725		4362		4362	1.5		0.0				0.4		0.1		0.0		0.0				2.7		0.1		5.2		2.1		0.3		0.0		1.0		0.0		0.1		0.0		30.2		0.0		0.0	4.2		0.0		6.9		77.9		120.8		8.2		416.2		32.8		359.9		21.6		89.1		2.6		82.1		19.1		206.4		0.0		59.9		0.0		0.0		0.0		115.6		0.0		0.0	22.6		10.6		278.6		28.6	0.0	32.5		0.0	0.0	0.0	0.0		0.0	180.4		17.5		4767.8		587.0		0.0	0.0		134.8		0.0		285.1	0.0		0.0		0.0		0.0	0.0		0.0