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Agnini, Claudia; Fornaciari, Eliana; Raffi, Isabella; Rio, Domenico; Röhl, Ursula; Westerhold, Thomas (2007): Age estimations of calcareous nannofossil biohorizons of the middle Paleocene to early Eocene at ODP Site 208-1262 (Table 1, Appendix B) [dataset]. PANGAEA, https://doi.org/10.1594/PANGAEA.691457, Supplement to: Agnini, C et al. (2007): High-resolution nannofossil biochronology of middle Paleocene to early Eocene at ODP Site 1262: Implications for calcareous nannoplankton evolution. Marine Micropaleontology, 64(3-4), 215-248, https://doi.org/10.1016/j.marmicro.2007.05.003

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Abstract:
Over the last several decades debates on the 'tempo and mode' of evolution have centered on the question whether morphological evolution preferentially occurs gradually or punctuated, i.e., with long periods of stasis alternating with short periods of rapid morphological change and generation of new species. Another major debate is focused on the question whether long-term evolution is driven by, or at least strongly influenced by changes in the environment, or by interaction with other life forms. Microfossils offer a unique opportunity to obtain the large datasets as well as the precision in dating of subsequent samples to study both these questions.We present high-resolution analyses of selected calcareous nannofossils from the deep-sea section recovered at ODP Site 1262 (Leg 208) in the South-eastern Atlantic. The studied section encompasses nannofossil Zones NP4–NP12 (equivalent to CP3–CP10) and Chrons C27r–C24n. We document more than 70 biohorizons occurring over an about 10 Myr time interval, (~62.5 Ma to ~52.5 Ma), and discuss their reliability and reproducibility with respect to previous data, thus providing an improved biostratigraphic framework, which we relate to magnetostratigraphic information, and present for two possible options of a new Paleocene stratigraphic framework based on cyclostratigraphy. This new framework enabled us to tentatively reconstruct steps in the evolution of early Paleogene calcareous nannoplankton through documentation of transitional morphotypes between genera and/or species and of the phylogenetic relations between the genera Fasciculithus, Heliolithus, Discoasteroides and Discoaster, as well as between Rhomboaster and Tribrachiatus. The exceptional record provided by the continuous, composite sequence recovered at Walvis Ridge allows us to describe the mode of evolution among calcareous nannoplankton: new genera and/or new species usually originated through branching of lineages via gradual, but relatively rapid, morphological transitions, as documented by the presence of intermediate forms between the end-member ancestral and descendant forms.
Significant modifications in the calcareous nannofossil assemblages are often “related” to significant changes in environmental conditions, but the appearance of structural innovations and radiations within a single genus also occurred during “stable” environmental conditions. These lines of evidence suggest that nannoplankton evolution is not always directly triggered by stressed environmental conditions but could be also driven by endogenous biotic control.
Project(s):
Coverage:
Latitude: -27.185833 * Longitude: 1.577000
Date/Time Start: 2003-03-24T00:00:00 * Date/Time End: 2003-03-29T00:00:00
Minimum DEPTH, sediment/rock: 106.65 m * Maximum DEPTH, sediment/rock: 201.04 m
Event(s):
208-1262 * Latitude: -27.185833 * Longitude: 1.577000 * Date/Time Start: 2003-03-24T00:00:00 * Date/Time End: 2003-03-29T00:00:00 * Elevation: -4755.4 m * Penetration: 583.9 m * Recovery: 502.5 m * Location: Walvis Ridge, Southeast Atlantic Ocean * Campaign: Leg208 * Basis: Joides Resolution * Method/Device: Composite Core (COMPCORE) * Comment: 54 cores; 493.9 m cored; 90 m drilled; 101.7 % recovery
Comment:
Ages provided for three different solutions based on GPTS (Berggren et al., 1995; Lourens et al., 2004) and revised timescale (Westerhold et al., 2007 doi:10.1029/2006PA001322, Westerhold pers. communication)
Parameter(s):
#NameShort NameUnitPrincipal InvestigatorMethod/DeviceComment
DEPTH, sediment/rockDepth sedmGeocode
Depth, top/minDepth topmAgnini, Claudia
Depth, bottom/maxDepth botmAgnini, Claudia
Depth, compositeDepth compmcdAgnini, Claudia
Depth, composite topDepth comp topmcdAgnini, Claudia
Depth, composite bottomDepth comp botmcdAgnini, Claudia
Sample code/labelSample labelAgnini, ClaudiaDSDP/ODP/IODP sample designation
Sample code/label 2Sample label 2Agnini, ClaudiaDSDP/ODP/IODP sample designation
Ageprofile Datum DescriptionAgeprof dat desAgnini, ClaudiaLO = lowest occurrence, LCO = lowest common occurrence, LRO = lowest rare occurrence, HO = highest occurrence, HCO = highest common occurrence, CO = crossover
10 Nannofossil zoneNannos zoneAgnini, ClaudiaNannofossil Zone; Martini, 1971
11 Nannofossil zoneNannos zoneAgnini, ClaudiaNannofossil Zone; Okada & Bukry, 1980
12 ChronozoneChronozoneAgnini, ClaudiaNotation (Chron.% from TOP), GPTS (Cande and Kent, 1995; Lourens et al., 2004)
13 ChronozoneChronozoneAgnini, ClaudiaNotation (Chron. % from Top) Option 1
14 ChronozoneChronozoneAgnini, ClaudiaNotation (Chron. % from Top) Option 2
15 Age modelAge modelkaAgnini, ClaudiaGPTS (Cande and Kent, 1995; Lourens et al., 2004)
16 Age, minimum/youngAge minkaAgnini, ClaudiaOption 1
17 Age, minimum/youngAge minkaAgnini, ClaudiaOption 2
18 Age, maximum/oldAge maxkaAgnini, ClaudiaOption 1
19 Age, maximum/oldAge maxkaAgnini, ClaudiaOption 2
20 Age model, optionalAge model optkaAgnini, ClaudiaOption 1
21 Age model, optionalAge model optkaAgnini, ClaudiaOption 2
22 Age, errorAge e±Agnini, ClaudiaOption 1
23 Age, errorAge e±Agnini, ClaudiaOption 2
24 Age, relativeAge relkaAgnini, ClaudiaRelative age to the PETM, Option 1
25 Age, relativeAge relkaAgnini, ClaudiaRelative age to the PETM, Option 2
Size:
1676 data points

Data

Download dataset as tab-delimited text — use the following character encoding:


Depth sed [m]

Depth top [m]

Depth bot [m]

Depth comp [mcd]

Depth comp top [mcd]

Depth comp bot [mcd]

Sample label
(DSDP/ODP/IODP sample designation)

Sample label 2
(DSDP/ODP/IODP sample designation)

Ageprof dat des
(LO = lowest occurrence, LCO =...)
10 
Nannos zone
(Nannofossil Zone; Martini, 1971)
11 
Nannos zone
(Nannofossil Zone; Okada & Buk...)
12 
Chronozone
(Notation (Chron.% from TOP), ...)
13 
Chronozone
(Notation (Chron. % from Top) ...)
14 
Chronozone
(Notation (Chron. % from Top) ...)
15 
Age model [ka]
(GPTS (Cande and Kent, 1995; L...)
16 
Age min [ka]
(Option 1)
17 
Age min [ka]
(Option 2)
18 
Age max [ka]
(Option 1)
19 
Age max [ka]
(Option 2)
20 
Age model opt [ka]
(Option 1)
21 
Age model opt [ka]
(Option 2)
22 
Age e [±]
(Option 1)
23 
Age e [±]
(Option 2)
24 
Age rel [ka]
(Relative age to the PETM, Opt...)
25 
Age rel [ka]
(Relative age to the PETM, Opt...)
106.65106.10107.19106.65106.10107.19208-1262A-10H-3,149208-1262B-12H-1,651 Ellipsolithus HONP12CP10C24n.1n?52374
107.67107.49107.85107.67107.49107.85208-1262B-12H-1,95208-1262B-12H-1,1312 Chiprhagmalithus LCONP12CP10C24n.1n?52525
107.67107.49107.85107.67107.49107.85208-1262B-12H-1,95208-1262B-12H-1,1313 Discoaster lodoensis LO or LO2NP12CP10C24n.1n?52525
110.09109.98110.19110.09109.98110.19208-1262B-12H-3,44208-1262B-12H-3,654 Girgisia gammation LONP11CP9bC24n.r?52796
113.52113.41113.62113.52113.41113.62208-1262A-11H-2,86208-1262A-11H-2,1075 Discoaster lodoensis LTO or LO1NP11CP9bC24n.3n531115322753603532565363253242536181515-2288-2312
113.52113.41113.62113.52113.41113.62208-1262A-11H-2,86208-1262A-11H-2,1076 Discoaster multiradiatus HONP11CP9bC24n.3n531115322753603532565363253242536181515-2288-2312
117.30117.04117.56117.30117.04117.56208-1262A-11H-4,149208-1262B-13H-2,27 Sphenolithus villae LONP11CP9bC24r.035C24r.063C24r.056534355369754073537505412753727541042627-1803-1826
117.30117.04117.56117.30117.04117.56208-1262A-11H-4,149208-1262B-13H-2,28 Sphenolithus orphanknollensis LONP11CP9bC24r.035C24r.063C24r.056534355369754073537505412753727541042627-1803-1826
118.09117.98118.19118.09117.98118.19208-1262B-13H-2,44208-1262B-13H-2,659 Tribrachiatus contortus HONP11CP9bC24r.056C24r.085C24r.0785348953786541645380454183537955417399-1735-1757
118.72118.61118.82118.72118.61118.82208-1262B-13H-2,107208-1262B-13H-2,12810 Sphenolithus radians LONP10CP9aC24r.073C24r.102C24r.0965353253840542205385954238538495422999-1681-1701
119.38119.27119.48119.38119.27119.48208-1262B-13H-3,23208-1262B-13H-3,4411 Discoaster multiradiatus HCONP10CP9aC24r.090C24r.118C24r.1125357853891542715390654286538985427877-1632-1652
120.01119.90120.11120.01119.90120.11208-1262B-13H-3,86208-1262B-13H-3,10712 Sphenolithus editus LONP10CP9aC24r.107C24r.133C24r.1275362153937543175395354333539455432588-1585-1605
120.67120.56120.77120.67120.56120.77208-1262B-13H-4,2208-1262B-13H-4,2313 Tribrachiatus orthostylus LONP10CP9aC24r.125C24r.149C24r.1435366653987543675400354383539955437588-1535-1555
121.30121.19121.40121.30121.19121.40208-1262B-13H-4,65208-1262B-3H-4,8614 Tribrachiatus bramlettei HONP10CP9aC24r.142C24r.164C24r.1585370954034544145405054430540425442288-1488-1508
123.88123.77123.98123.88123.77123.98208-1262B-13H-6,23208-1262B-13H-6,4415 Tribrachiatus digitalis morphotype HONP10CP9aC24r.211C24r.222C24r.2165388654216545965423154611542235460377-1307-1327
124.87124.76124.97124.87124.76124.97208-1262A-12H-3,65208-1262A-12H-3,8616 Tribrachiatus bramlettei decreaseNP10CP9aC24r.237C24r.244C24r.2385395454285546655430054680542925467277-1238-1258
125.08124.97125.18125.08124.97125.18208-1262A-12H-3,65208-1262A-12H-3,10717 Tribrachiatus digitalis morphotype LONP10CP9aC24r.243C24r.249C24r.2435396854300546805431454694543075468777-1223-1243
125.50125.39125.60125.50125.39125.60208-1262A-12H-3,149208-1262A-12H-4,218 Tribrachiatus contortus LONP10CP9aC24r.254C24r.258C24r.2525399754329547095434354723543365471677-1194-1214
127.45127.34127.55127.45127.34127.55208-1262A-12H-5,23208-1262A-12H-5,4419 Discoaster diastypus LONP10CP9aC24r.307C24r.303C24r.2975413154469548495448554865544775485788-1053-1073
133.34133.23133.44133.34133.23133.44208-1262B-14H-5,44208-1262B-14H-5,6520 Tribrachiatus bramlettei LONP10CP8bC24r.464C24r.446C24r.4425453554916552995493255316549245530888-606-622
135.87135.76135.97135.87135.76135.97208-1262A-13H-3,65208-1262A-13H-3,8621 Fasciculithus HONP10CP8bC24r.532C24r.505C24r.4995470855101554815511755494551095548887-421-442
139.72139.71139.72139.72139.71139.72208-1262B-15H-3,28-29208-1262B-15H-3,29-3022 Rhomboaster calcitrapa gr. HONP10CP8bC24r.636C24r.616C24r.6185497255454558565545655858554555585711-75-73
139.80139.78139.81139.80139.78139.81208-1262B-15H-3,35-36208-1262B-15H-3,38-3923 Fasciculithus / Zygrhablithus CONP10CP8bC24r.638C24r.620C24r.6225497855464558685546855874554665587123-64-59
139.80139.78139.81139.80139.78139.81208-1262B-15H-3,35-36208-1262B-15H-3,38-3924 Fasciculithus decreaseNP10CP8bC24r.638C24r.620C24r.6225497855464558685546855874554665587123-64-59
139.82139.81139.82139.82139.81139.82208-1262B-15H-3,38-39208-1262B-15H-3,39-4025 Zygrhablithus bijugatus LCONP10CP8bC24r.638C24r.621C24r.6245497955468558745547055875554695587511-61-55
139.92139.84140.00139.92139.84140.00208-1262B-15H-3,41-42208-1262B-15H-3,57-5826 Campylosphaera eodela LCONP10CP8bC24r.641C24r.625C24r.629549865547255879554945590755483558931114-47-37
139.99139.97140.00139.99139.97140.00208-1262B-15H-3,54-55208-1262B-15H-3,57-5827 Tribrachiatus bramlettei LRONP10CP8bC24r.643C24r.628C24r.6335499155490559025549455907554925590423-38-26
140.02140.01140.02140.02140.01140.02208-1262B-15H-3,58-59208-1262B-15H-3,59-6028 Rhomboaster LO/R. calcitrapa gr. LONP9bCP8bC24r.644C24r.629C24r.6355499355496559095549755910554965590911-34-21
140.15140.15140.15140.15140.15140.15208-1262B-15H-3,72-73208-1262B-15H-3,72-7330 Fasciculithus alanii HONP9aCP7C24r.647C24r.642C24r.6425500255514559325551555933555145593211-162
140.15140.15140.15140.15140.15140.15208-1262B-15H-3,72-73208-1262B-15H-3,72-7331 Fasciculithus decrease in diversityNP9aCP7C24r.647C24r.642C24r.6425500255514559325551555933555145593211-162
148.19148.08148.29148.19148.08148.29208-1262Añ14Hñ4,85-86208-1262Añ14Hñ4,106-10732 Campylosphaera eodela LONP9aCP7C24r.863C24r.868C24r.8635555356232566135624956629562415662188711691
149.90149.79150.00149.90149.79150.00208-1262A-14H-5,106-107208-1262A-14H-5,127-12833 Zygrhablithus bijugatus LONP9aCP7C24r.908C24r.912C24r.90755670563695674856387567685637856758910848828
151.82151.71151.92151.82151.71151.92208-1262B-16H-4,45-45.1208-1262B-16H-4,66-66.134 Fasciculithus alanii LONP9aCP7C24r.960C24r.963C24r.95955802565295691456546569315653756922981007992
153.32153.21153.42153.32153.21153.42208-1262A-16H-5,45-45.1208-1262A-16H-,566-66.135 Ericsonia robusta HONP9aCP7C25n.002C25n.011C25n.011559055665157046566695706456660570559911301125
154.16154.05154.26154.16154.05154.26208-1262B-16H-5,129-129.1208-1262B-16H-5,149.9-15036 Fasciculithus richardii gr. LONP9aCP7C25n.146C25n.134C25n.139559755672057120567355713556727571277711971197
154.61154.50154.71154.61154.50154.71208-1262B-16H-6,24-24.1208-1262B-16H-6,45-45.137 Discoaster multiradiatus LONP9aCP7C25n.224C25n.193C25n.195560135675257152567675716756759571597712291229
155.66155.55155.76155.66155.55155.76208-1262A-15H-2,89208-1262A-15H-2,89-11038 Discoaster multiradiatus LRONP8CP7C25n.406C25n.331C25n.328561025682657226568425724356834572348813041304
155.87155.76155.97155.87155.76155.97208-1262A-15H-2,110-110.5208-1262A-15H-2,131-131.539 Discoaster okadai HONP8CP7C25n.443C25n.359C25n.358561205684257243568575726056850572518813201321
156.08155.97156.18156.08155.97156.18208-1262A-15H-2,131-131.5208-1262A-15H-3,2-2.540 Discoaster delicatus gr. LCONP8CP7C25n.482C25n.388C25n.388561395685757260568735727756865572688813351338
156.92156.81157.02156.92156.81157.02208-1262A-15H-3,65-65.5208-1262A-15H-3,86-86.541 Discoaster delicatus gr. LONP8CP7C25n.625C25n.504C25n.508562085692057327569365734456928573368813981406
157.13157.02157.24157.13157.02157.24208-1262A-15H-3,86-86.5208-1262A-15H-3,107-107.542 Discoaster okadai LONP8CP7C25n.662C25n.536C25n.54156226569365734456957573675694557355111114151425
157.35157.24157.45157.35157.24157.45208-1262A-15H-3,107-107.5208-1262A-15H-3,128-128.543 Discoaster nobilis gr. LONP8CP7C25n.699C25n.578C25n.58456245569575736756980573915696857379111214381449
157.89157.68158.10157.89157.68158.10208-1262A-15H-4,2-2.5208-1262A-15H-4,44-44.544 Fasciculithus tonii LONP8CP6C25n.812C25n.685C25n.69356299570045741757049574655702757441222414971511
158.00157.89158.10158.00157.89158.10208-1262A-15H-4,23-23.5208-1262A-15H-4,44-44.545 Ericsonia robusta LONP8CP6C25n.794C25n.706C25n.71456291570275744157049574655703857453111215081523
158.21158.10158.31158.21158.10158.31208-1262A-15H-4,44-44.5208-1262A-15H-4,65-65.546 Discoaster sp.1 HCONP8CP6C25n.848C25n.747C25n.75756317570495746557072574885706057476111215301546
159.16159.15159.17159.16159.15159.17208-1262A-15H-5,2-2.5208-1262A-15H-4,149-149.547 Discoasteroides megastypus LONP8CP6C25r.016C25n.929C25n.934563995715857576571605757857159575771116291647
159.49159.38159.59159.49159.38159.59208-1262A-16H-5,22-23208-1262A-16H-5,43-4448 Fasciculithus richardii gr. LRONP8CP6C25r.033C25n.989C25n.99056429571815759857202576195719257608111016621678
165.00164.79165.21165.00164.79165.21208-1262A-16H-1,133-134208-1262A-16H-2,25-2649 Discoaster sp.1 LONP8CP6C25r.481C25r.332C25r.33656950576435805157676580905765958071161921292141
168.05167.94168.15168.05167.94168.15208-1262A-16H-3,148-149208-1262A-16H-4,19-2051 Heliolithus bukryi HONP7CP6C25r.728C25r.619C25r.61957237580445844058076584715806058455161525302525
169.52169.41169.62169.52169.41169.62208-1262A-16H-4,145-146208-1262A-16H-5,16-1752 Sphenolithus anarrhopus gr. LCONP7CP6C25r.847C25r.768C25r.76057376582535863458282586625826758647141427372717
169.52169.41169.62169.52169.41169.62208-1262A-16H-4,145-146208-1262A-16H-5,16-1753 Sphenolithus sp.1 LCONP7CP6C25r.847C25r.768C25r.76057376582535863458282586625826758647141427372717
169.95169.84170.05169.95169.84170.05208-1262A-16H-5,37-38208-1262A-16H-5,58-5954 Heliolithus kleinpellii HCONP7CP6C25r.882C25r.811C25r.80457417583125869358342587235832758708151527972778
170.15170.05170.25170.15170.05170.25208-1262A-16H-5,58-59208-1262A-16H-5,79-8055 Heliolithus bukryi LONP7CP6C25r.899C25r.832C25r.82657436583425872358370587515835658737141428262807
171.50171.39171.60171.50171.39171.60208-1262B-18H-3,40-41208-1262B-18H-3,61-6256 Discoaster LO/D. mohleri LONP7CP6C26n.054C25r.968C25r.96757573585295891358568589525854558929202030152999
171.71171.60171.81171.71171.60171.81208-1262B-18H-3,61-62208-1262B-18H-3,82-8357 Sphenolithus cf. conicus LONP6CP5C26n.169C26n.004C26n.00357614585685895258614589985859158975232330613045
171.92171.81172.02171.92171.81172.02208-1262B-18H-3,82-83208-1262B-18H-3,103-10458 Heliolithus cantabriae HONP6CP5C26n.285C26n.171C26n.16557656586145899858655590395863659020212131063090
172.13172.02172.23172.13172.02172.23208-1262B-18H-3,103-104208-1262B-18H-3,124-12559 Fasciculithus clinatus LONP6CP5C26n.401C26n.297C26n.28957697586555903958687590725867159055161631413125
173.22173.16173.27173.22173.16173.27208-1262B-18H-4,66-67208-1262B-18H-4,78-7960 Ellipsolithus LCONP6CP5C26n.998C26n.925C26n.91557910588305921858854592465884259232121433123302
173.93173.88173.97173.93173.88173.97208-1262B-18H-4,138-139208-1262B-18H-4,148-14961 Discoasteroides bramlettei gr. LONP6CP5C26r.032C26r.046C26r.05358009589865939759005594195899559408101134653478
174.05173.97174.12174.05173.97174.12208-1262B-18H-4,148-149208-1262B-18H-5,13-1462 Heliolithus kleinpellii LONP6CP5C26r.038C26r.055C26r.06358025590055941959037594565902159438161934913508
174.44174.34174.54174.44174.34174.54208-1262B-18H-5,34-35208-1262B-18H-5,55-5663 Sphenolithus anarrhopus LONP5CP4C26r.056C26r.084C26r.09758079590855951159128595605910759536212535773606
175.18175.07175.28175.18175.07175.28208-1262A-17H-2,118-119208-1262A-17H-2,139-14064 cf. Discoaster LONP5CP4C26r.089C26r.135C26r.15158179592295966559280597195925459692262737243762
175.39175.28175.49175.39175.28175.49208-1262A-17H-2,139-140208-1262A-17H-3,10-1166 Sphenolithus s p.1 LONP5CP4C26r.099C26r.152C26r.16958208592805971959323597625930659746222137763816
175.81175.70175.91175.81175.70175.91208-1262A-17H-3,31-32208-1262A-17H-3,52-5365 Heliolithus cantabriae LONP5CP4C26r.118C26r.174C26r.18858265593545978759384598125936959799151338393869
184.91184.83184.99184.91184.83184.99208-1262B-19H-5,34-35208-1262B-19H-5,50-5167 Fasciculithus billii LONP5CP4C26r.504C26r.549C26r.549594266045260840604686085660460608488849304918
188.03187.92188.13188.03187.92188.13208-1262C-10H-4,50208-1262C-10H-4,7168 Fasciculithus tympaniformis gr. LONP5CP4C26r.672C26r.700C26r.69659934608866125960915612876090261274141453725344
190.76190.65190.86190.76190.65190.86208-1262C-10H-6,23208-1262C-10H-6,4469 Fasciculithus pileatus LONP4CP3C26r.796C26r.811C26r.81060306612116159061236616156122461602121356945672
190.76190.65190.86190.76190.65190.86208-1262C-10H-6,23208-1262C-10H-6,4470 Fasciculithus LO / F. ulii LONP4CP3C26r.796C26r.811C26r.81060306612116159061236616156122461602121356945672
191.72191.51191.92191.72191.51191.92208-1262B-20H-2,77208-1262B-20H-2,11971 Neochiastozygus perfectus LONP4CP3C26r.840C26r.851C26r.85060437613146169461364617446133961719252558095789
193.92193.82194.02193.92193.82194.02208-1262B-20H-4,8208-1262B-20H-4,2972 Sphenolithus LONP4CP3C26r.940C26r.938C26r.93860738615816196161604619846159361973111160636043
196.85196.74196.95196.85196.74196.95208-1262C-11H-2,122208-1262C-11H-2,14374 Chiamolithus bidens gr. LONP4CP3C27n.638C27n.569C27n.56961147619356231061961623356194862322131364186392
201.04200.94201.15201.04200.94201.15208-1262C-11H-3,92208-1262C-11H-3,11375 Toweius pertusus LONP4CP3C27r.488C27r.444C27r.44961873625156289562544629276253062911151670006981